Tesis Doctoral presentada por JORGE LUIS RUIZ RAMÍREZ IRTA !"#" $%&'%( (!"!)' $ &%*% &%* !)$#% $! +, #& &#%)!" %( - / 0 1 . - ! $ "# % & % ) 4, 3 ( * + ,$ - . ' . 1 . , , / , / ., , , 0 + ( 2 3 ( . 5 , , 5 , , 5 6 60 6 60 0# 5 !+ +8 # 0# 7 / / "+ 7 / + 6 6 0 # 5 + 7 ) / / !+ +8 0 , " 0 9: -0 9 9 ; ;07 09 4 9 ; * 9* 9 = ; *0 <: ;-: => ;? * = *9 : ;: ;- 4 ;-: 5 2 ? = 0 ) ; 8 . 3 =2 3 . : , A . B A . 5 . B 5 . ? 7 ) , , , 5 * / , / + 1 + + . , 0# , !+ +4 " , + * " , GI8GJK# ) A ! " / , ( , , = ! 5 8+ +" , , # 1 & 2 , = 8+ + K A # ; 0 0# + ,B , A , -FGGL8GKMIF 0 + , , ) !* / , 2 ;, A ) + ) ( B / , , / + /5 -FGGG8GHIJ8 K8GK ) 1 E"# ) , 3 + , + 5 # , A !D !C9* + " , 5 ) A 3 . B ' # + , , . / = + , + 3 # # 7 ) C , 0 , , , 3 , , , ) , 5 , , 0 # , ) 7 ) # , , , 5 , B , 2 3 / , @ , # ; = / B * , # B + = ) , + * / : 0 ) = / $ N :O 2 B 5 B = 5 - + # , A ( " , ( 7 ) ? , 3 + , , / @ ) 3 / N 0 C P , A/ # , A C , 7 ) / B) ., 7 , 5 , A/ # =3 + 3 ) , )) 1 B , + , , A/ # = 1 ) 1 # , 3 A 9* # B 3 , / , , 5 9* ) 1( 9* ) , A B 3 , / 1 3 ) 3 /, , 3 / . , , A/ # 4 , 3 5 , ) # D @ / # , A , 3 ) 1 !7 , ; )) A. 7 ) : @ :2 1 . , P 5 = @ ) 4 + # :2 1 . , = = 9 ) C , /& 4 , = # , A/ = A. :2 , . , = + = / 8 / / 909: = 0;+ = ;* 9E ( >*0 +; 0; +9* ;* 09 + ################################################################################################### 0 ################################################################################################## ) ) ######################################################################################### > + ################################################################################### - , - C , - ) 5 ####################################### ##################################### ) ) ################################################################################ ####################################################################################################### ###################################################################################################### ####################################################################################################### ######################################################################################################### I ############################################################################### L ################################################################################### K I# F#I# 9 ( ) F#F# 9 ( ) ################################################################ K , /5 ######################################################### H 25 ######################################################### H 3 !7#"#4) ,#, #%8(9:#& ########################################################## Q L# 2'+ % ) ) ,6!'#7%" ###################################################### 2'+ % 3 )'(%$+&&#4) 5 F# 9 ( ) 5 I# 0 , ( I#I# 0 5 I#F# ; I#L# ########################################################################### II , ################################################### II ###################################################### IF ################################## IF ########################################### IK ######################################## IH ################################################################################ IH ############################################################################### IJ ( 5 ( F#I# : , , F#L# : F#K# II , F# F#F# ################################################### , F#H# : F#M# ; ) , 8= ########################################################## IR ########################################################################### FG @ L# ( L#I# C ############################################################ 3 ################### FL ####################################################### FL ############################################################# FL , #### FK L#I#I#L# ,$ ################################################################################## FK L#I#I#K# #################################################### FH ########################################################### FM ############################################################ FJ L#I#I# = 5 ( FI , . L#I#I#I# 9 B L#I#I#F# + , / L#I#I#H# + L#I#F# L#I#F#I# , 1 L#I#F#F# + L#I#F#L# L#F# ) , , ) ################################# , ( FQ LL ################# LK ########################################################################### LK L#F#F# S , L#L# =B ### ######################################################## , L#F#I# ; A FJ . LH ########## LM ###################################################### LM ################################################ LM ######################################################## LM ######################################################## LM , L#L#I# =B L#L#I#I# =B ################################################## 5 L#L#I#F# =B ,/ L#L#I#L# =B ) L#L#I#K# =B 3 / L#L#I#H# =B 5 L#L#I#M# =B , ############################## LJ ##################################################### KG ) L#L#F# =B ### KG ######################################################### KG ################################################################################################### KL L#K# K# 5 2'+ % I# F# ; , @ 25 3 ###################################################################### KK ;'%$%" ######################################################### HH ############################################################################# HJ I ######################################################################################### HR '!(# !" 5 L# ; , K# 0 F ######################################################################################### A L K ##################################### ML K#I ; , L ################################################################################ MH K#F ; , K ################################################################################ MQ ################################################################################## JI 25 JF H# 2 M# 5 , , MG / 2'+ % 3 !<'+(! ###################################################################### ( *!'!(" %: $(5=&+(!$ > * *? " * !" $(#!$ ' $#::!(!)' !7! " " :+)&'#%) %: @ '!( &'#7#'5 )$ @ '!(&%)'!)' T################################TTTTTTTTTTTTTT# JL ####################################################################################################### JH ############################################################################################# JM I# F# = 1 ############################################################################ JJ ############################################################################ JQ ################################################################################# RL ############################################################################################### RK L# K# U H# 5 V 2'+ % 3 (%:# !" %: @ '!(&%)'!)' @ '!( &'#7#'5 )$ '!<'+(! #) &(+"'!$ $(5=&+(!$ %#) )$ #) )%)=&(+"'!$ $(5=&+(!$ %#) T###TT##### RJ TTTTTTTTTTTTTTTTTTTTTTTTT### RQ ############################################################################################# QG I# F# = 1 ############################################################################ QI ############################################################################ QL TTTTTTTTTTTTTTTTTTTTTTT# QQ TTTTTTTTTTTTTTTTTTTT# QQ TTTTTTTTTTTTTTTTTTTTTTT### QQ L# K# + H# U M# 5 V 2'+ % F# = L# ! '#%)"># ,!'@!!) @ '!(&%)'!)' &%)'!)' ( *!'!(" #) $(5=&+(!$ *+"& !" T###TTTTTTT##### IGL ####################################################################################################### IGH ############################################################################################# IGM )$ '!<'+(! I# 3 1 ############################################################################ IGJ ############################################################################ IGQ @ K# + H# U M# 5 TTTTTTTTTTTTTTTTTTTTTTT# IIR TTTTTTTTTTTTTTTTTTTT# IFG TTTTTTTTTTTTTTTTTTT###########TT IFG V 2'+ % 3 ::!&' %: A )$ ,!'@!!) @ '!(&%)'!)' )$ '!<'+(! ####################################################################################################### IFJ ############################################################################################# IFR 1 ############################################################################ IFQ ############################################################################ ILF ############################################################################################# IKF ################################################################################# IKF ############################################################################################### IKF $! %" (!"+ ' $%" ################################## IKJ ################################################################################## IKQ L# K# + M# 5 )$ IFH F# = U ( *!'!(" #) *+"& !" #) $(5=&+(!$ > * TTTTTTTTTT# I# H# &%)'!)' %) '>! (! '#%)"># V 2'+ % 3 #"&+"#4) 8 %, I# ) , 2 F# ;5 ' ,$:=FK , 2 ############################################################################ L# ;5 * + , 2 K# ;5 A ########################################### / , 2 H# 5 2'+ % @ IHM , ### IMI ####################################################################### IMK 3 %)& +"#%)!" ######################################################################## IMJ %)8(!"%" #################################################### IJI : ,;'#&% ####################################################################################### FGQ 25 )!<%"3 %*+)#& &#%)!" .)$#&! IHK I# + , , F# + , 3 / ############################################################### ' /, /5 ############################################################################################ L# + K# 5 ( 2 ! ####################################################### , 2 " LR ############################################################################### LQ B 0 J# ;3 R# 0 . , 2 . ( , A , . L K, ' ################### KI ####################### HJ ! " ########################################################## Q# + , IG# = ( ################################## 2 , , II# = IF ############## H# M# IF 5 / ( IF !W0#;" , 1 ) 5 ML MM . ##### 3 / '8 ( ####### IKM IHR @ C I# ) , ######################################################## C F# C L C K# C H# C M# 25 , 2 J# ' R# C Q# C II# HQ ################################################################ MI , , A , + 3 . 5 , . ,/, ' #################################################### MH ####### MM ################################################# MJ #################################### MQ #################################################### JG , 2 , 25 A ! C" ( IF# !U ) SU " . ################################################################ C IL# ) C IK# ) C IH# 1 ) IHG ##################### IHI ####################### IHF , 2 ' ( ' ############################################################ IM# . , ' C MK " , !:=" C MF , + ' C F L K ###### L K ########################################################## , IG# I ################### ;3 !, C LJ . ( C ########### . , C , 5 LI 1 ( IJ# ,$:=FK ' ' ,$:=FK ################################# 1 ) ( IHH , 1 IHJ ' ############### IHR @ C IR# . , ,$:=FK * + ' C 1 * + C ' IHQ ) ' , 2 , , ,$:=FK , FG# / @ 1 ' ########################################## IQ# , 1 ( ########### IMG ' ############################# IML 9 + 5 955 95 +1 & / / ! " ) C =' 0C0 0 UC ; ;, 0 C : " . 5 $ !9 . $ ) > :9 9 . , U 5 ; : . . !9 . 8 " X X 5 -; - ; -*; - * ; = = =: = * * * + + ** * * * ,$ 8 ,$:=F ,$ ,$:=FK ,$ ,$:= ,$ :; . : * Y$Z[ F1 ' FK 1 ' : 5 ; ) ! 2 ; ) " @@ F + 5 0 =:; : 0 ) !0 ) ; 2 " , := ) , =' 5 P P ! 2 5 " , !U . + SU " + , !U SU 8U * +" + , U @@ !X SU " 8U * +8 ; A 5 ) 5 A , ( ) , ! #; " ) . ' + / ! : 2 ) ) ( , ( ; , ) 0 ) 3 ,/ ,/ , 3 , % , . # , ) ( ,/ #; , + "# ,/ ) ) ,$ 5 25 3 , 5 , , B , , , # ; ,/ , / ) , , , 2 , 2 ( , , ; , S !; ,# F"# ; 3 , ,/ 5 ,$:= , , 2 # , , , ,/ ' 5 , 2 ( 5 ) \ , 5 ,/ 5 * + ,/ ,/ ) \ ' !; ,# I" , A ) ) # - , ) , , 5 ,$FK ,$ * + , 2 * + @@ , 2 ( ; ,/ 5 1 ' # , ( , , ,/ # B 1 , 3 , , 0 , # + ( , , # ( @@ ) FGGH - , 0 , 2 , ( % * + ,$ ) ! " , , #0 1 . / , ( ( 3 , 5 , 3 , # ; 1 , I ) ) ' #; , " , 5 L * + K ) A ( 2 , , KH FRQ / . 1 / ! , ! , ( ! ( ( L" #- ) 5 5 ) K"# 5 3 # 5 ,$ ( ' ) ' . , , 3 ) . , , 5 1 ) / #; ) / , 1 ##- GJ ( KH ) . GM ' 5 ) ' 5 . F , 2 , , , , ) , ) " , 5 , ( 5 !- , , 2 ( 5 ) A ) . 3 ' , , # 3 , ( - S ' . ! ! " , , 2 , ) ) 5 ) ( @@ ' 3 , ' ! ) ' # ) # - A , 1 / , 5 , 2 , ) @@ ( ( ,$:=FK ] / , 3 , 5 / / ) 3 ( # , 1 , 5 GM, ' , ( ! 5 1 ) ( A ( ( ) ) . , 2 . ) 1 * + ,$:=FK ^ M F# , ) ]GJ . #- * + , , ! ] G GH" HJ, , , 2 # - * + 5 ' ' ) #* 1 " #; ( . 3 5 S ) , , ) " , ) GR, # / 1 5 1 , 1 , 0 , 1 %V ) 1, V ) 1 ! " V 8, S 8 , #* , ) , 55 , I1 1 ) , 5 , 1 ( 1, 5 1 * + 5 ! , KH 8 #$ V ) 5 V , 5 1 8 55 ) , 1 8 5 1 1 , ) , ) 8 1 5 V 11 FRQ 8 V1 S SU 5 ) 5 1 1 3 V 5 # 1 1 # 1 ) 8 # 1 ! ( # 1 8 5 1 1 8 1 5 1 8, ,, ) K1 1 5 1 U L G#M U V 5 1 1 55 5 # $ ) G#J 1 1 , V V V 55 L" 1, 1 ) ,$ ) ! , K"# 1 1 #C # 5 1 1 5 !1 1 8 # 1 V , , 8 1 V 5 # 1 V 8 , 55 1 8 1 1 , 5 1 1 # , ) ) # C 1, 55 F1 1, 5 1 1 3 " 5 1 5 1 8 5 V 1, , , # 1 5 5 1 ) 1 V 1 1 , 1 5 , ( 1 V ) V 1 1 ,$ 8 V U V ! " * + 5 , U V , 1, V1 1, 1 ) KH V V V 1 , 5 S 1 # 1 " " 8 V @@ , 8 , 1 1 # 1 1 8 1 1 1 1 # 1 5 8 ! ] G#GH" 1 V 11 1 1, * + , V1 1 ,$:=FK ^ M#F# 9 1 55 5 1 * + V , 1 V 11 1 V ) 1 1 8 1 5 ) ) 1 5 " 1 1 1 5 ,, 5 5 ) V 5 8 5 5 8 ) 1, 1 8 1 , 8 , ) 1 ) 1 5 5 G#R 5 1 ) # 9 1 1 1 V 1 1 ! 1 G#M 5 8 # 1 V 1 # V1 1 1 1 1 1 1 , , 1 # ) 1 1 1 , 1 V 1 55 , , @@ 1 @]G#J# 1 * + ,$:=FK ] H#J 3 1 1 55 1 1 5 ) 8 , ) 1 # 1 ; ( _3 , 0 , * + ,$ % N ) _ ! " , , , ' . 3 , 3 ) # + 3 , 3 ) , # -_ , 5 ) , 3 , ( ! " ' _ # -_ , F) ! ) ' , ' O #; , _5 , 1 ) L" , _ L K) 5 ,$ 5 ' , KH FRQ ! , , K"# - ) 3 N 5 * + , `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ntroducción y Objetivos Capítulo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orge Ruiz Ramírez K , Tesis Doctoral 5 , * + , 5/ # ; , 5 ) * + , 3 ,$# , ) ' 1 # - , 2 5 . , , 1 / 5( , , 5 ) ) 8 , B 5 !C ; 2 , : " ; # FGGI"# 5 , 8 K#I#F !> , 5 , , , 3 , 1 # , 1 , 9 K#F#I , IFa + ( / B, : , , 0 % B ! !C ) 5 3 1 , / , 8 , 5 : , ,$ 5 2 , S " ? IQQR"# , ?B? , ,6!'#7% 8!)!( 8 ,# % ( 0 1 3 , 2 , 5 , ! S ) 3 , . * + ,$ , # , 2 , 5 ) S " , 2 , Introducción y Objetivos 2 Capítulo I # ? ? C? D = ,6!'#7%" !" !&2:#&%" - ( ) , /5 ( ) , 5 B 1 ) : 1 , % % # + ) # 1 5 # # # !FGGK"# 5 . 5 # # MR KH8HI# # !IQQI"# $ , ( I# 0 ) # , 7 ) ! . 1 " ; , A# # !IQQR"# , ) ( , 8 #; ( % 2 # ' $ % ! && ( #$)# # IFR8IKK# # -# # !IQQJ"# ;55 5 , 1 1 5 5 8 1 5 1#* ' $ + , JK IQL8IQR# # -# :2 +# !IQQR"# 1 55 5 1 ,$ * + 1 ,8 ) 1 8 5 8 1 #* ' $ + , JJ LRJ8LQF# A :# + ;# +# =# 0# =# 0# !FGGF"# , , U 8 # &-' ( #$)# # IFM8IFJ# 1 :# B # # + =# # +# = # !IQQK"# V 8 / F# 0 , 5 ) " 8 # L# 0 5 * + ,$:=FK , 1 ) ' KH / FRQ / # K# * + , . 1 ) ' ' ( 8 ( # . , C , 2 ( H # M Jorge Ruiz Ramírez 3 5 C 1 8 1 ## $ LJ FFQ8FKL# C # !FGGI"# ; : % 5 8 ) #." , / ( % 0 JH H8IG# C : # !IQQR"# , : # 9 FK * ) FGGL C : P % 1 ,%SS5 # S # , / 8 # & 8? 5 # , # - 3 + =# !FGGG"# ) 5 1 , 0 $ - :, 1 8 1 5 5 ) # # $ HM I8M# / 8 # & 8? 5 # , # - 3 + =# !IQQQ"# : , , 5 :, 1 8 1 5 5 ) 8 #+ ' MJ KFL8KFJ# / 8 # / 8 # & 8? 5 # , # - 3 + =# !FGGK"# 8 1, V ,$ 5 8 1 3 # # $ MJ MFH8MLF# # + , # !FGGF"# 2 , , 8 ( #% IGH RH8QM# # -# # !IQQH"# : 55 8 1 1 # # $ KG FI8LI# -# # # 0# # !FGGG"# ;5 8 ( , , # (( $ ( Tesis Doctoral * / # 8 # QK8QK# -# # # # !IQQM"# : 5 1 ,1 8 1 1 5 8 1 1 , , # * ' $ + , JG HFM8HLG# -# # # # !IQQQ"# 1 5 5 ,$ 1 , , 5 8 1 # # $ HF FMJ8FJL# = -# :2 1 . # =# : ( 2 # - , . *# !FGGL"# 5 5 V 5 3 5 1 # % (( # * / # +2 8; , A ! = / -# 3 # + # =# # !IQQR"# ;55 5 , , , 5 1 # # $ KQ IKH8IHL# = U V . $# # !IQRJ"# + ) , 8 . 5 5 # $# # = U V . !; #" C # * V 4 U% = 0 UU # LLH 9 ) =# # # , =# 0 # # * #-# # !IQQK"# + , 5 5) , 5 , # # 5 1 3 1 8 5 1 # 1" 2 $ KG IJQ8IRH# ) # +# =# ) *# !IQRR"# + . , ,, , # ( " ML KH8KQ# # # : 1) 8 Introducción y Objetivos . ,, V , +# !IQQK"# 1, 1 , ) , 8 1 5 5 ) ## $ LR IIJ8IFF# . # #+ ) # =# / +# !IQQR"# : 1 5 1 % 5 5 , #+ . '( LI HL8HR# .8+ # # + ) # B # / +# !FGGG"# ,, 5 8 1 55 5 5 , # + . '( LL QI8QH# : 1 ) . ,, +# 0 =# * + -# # .. -# # !FGGF"# 5 V , , 1 # # $ MG JJ8 RL# Capítulo I :. . U J # !IQMR"# + V ( ) # + ) ' FF QRI8QRM# :. . U # !FGGF"# , , #+ 3 2 IL FIH8FFH# :. . U # X 0# !IQML"# + V 5 1 5 # + ) ' IJ JK8JR# # C =# C . :# 2 C# !IQQQ"# 3 55 , , , U V 5 8 1 # # $ HI FHH8FMG# # # : 1 ) . ,, +# : +# =# !IQQH"# : 3 5 8 1 55 8 1 , ) , # * + $ MG IIRL8IIRM# . D Revisión Bibliográfica Capítulo II 2 B? , # ( , 8 1 1 , !IQQR" , B?B? !:#)#&#4) $! 6 *4) &+( $% II B .2 . ) 8 , ; ( 2 , , , , , 2 # < 5 , , ' . . , !, 2 " , , , 8 , ' # ; + ;, A 5 , 4 4 ( 2 4 4 / 5 3 8, 8 4 4 5 4 , 5 ) 4 8 8 2 !: - ( , 5 ) 2 , / 8 IQRF"# IQMJ"# 3 6 ( ! - 4 4 , ) 4 8 / ) 6 4 1 ( , 4 6 ) , 4 4 5 , ) 4 4 , 3 4 8 . , B 5 5 ' 1 5B , , 3 , # B? ? "'(+&'+( 5 %* %"#&#4) , ) , , . , , , ) 8 . 8 8 # , Jorge Ruiz Ramírez IF ) Tesis Doctoral ' !IQQL"# ; , ) I# + , ) ! , L# + IL Qa 2 5 *%)!" MR Fa =' ( !8E) (%&!$!)&# L Ra !($% , )&% R Ja $ )B . !IQQL"# 3 5 b) , 8 3 / ' - V !IQRR" , , ' !8E) &(+&!" 8!);'#&%" !-Vc-"c0 !-Vc-"c-Vc-c0 -Vc -Vc0 = ) /5 L"# ( /, !($% #,;(#&% # * )& ( c0 FH8JHa 0 +! 7 F# + /, ' , 3 / /5 + = JH GGa / + IQ GGa * , I MHa -/, F HGa = G MHa + 1 - V !IQRR"# 9&!(!" = 2 1 . $ 6%F ? 5 . + C : I FGa B?C? # %" $! 6 *4) &+( $% -V% \ ; ) , ( P1 \ -% % B #= \ 0% 0 !IQQM"# \ % , 8 ? " ( , " ( , B # ; ' B Revisión Bibliográfica ( Capítulo II , 1 , , , # , . ) 1 8 8 3 8 , , . 5 , B A , , , . - 3 8 3 , , % 8 , # F#8 0 . 25 8 , 8 / #; 5 , 5/ , / # # 5 ( 2 , % , S , !3 5 - 8 , , , 8 "# 3 , . , ( 5 3 , 5 3 , / / , 2 8 , /, 2, C # IQQJ\ . 5 , 2 1 2 C / # ; 8 5 ) 8 8 # , 8 # IQQL\ ( , 3 , 8 8 , S 8 # , , - ( , I#8 ; 5 / , ( % . , , 5 , / , ' # 1 , "# ; ! # IQQQ" 3 / , !, 8 2 # IQQL\ + IQQR\ , 5 1 ) ) , 3 / , , 5 ( 5 8 ) , 3 5 , 3 IL , , 3 , / 5 ( 8 Jorge Ruiz Ramírez IK , A Tesis Doctoral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evisión Bibliográfica Capítulo II , ! B . , ! , 3 5 " ) B 5 8 .' , # ; " ,2 1 # , ? ? IH (! ( &#4) $! * + * '!(# 5 # 2 2 , ( (#* / 5 , , B 1 , 5 ( . ; , 8 ! "# 5 ?C?B? ' , 5 3 , 3 8 ) / / 8 !$ 2 . , , , 5 ) , . IQQL"# /3 # - , , 8 .' 1 $% 2 - 5 , , , 8 ( ) ?C? 8 5 5 8 ( $ , IQQI" , , 2 KR 1 F 6+ ! , #; ( FK 3 5 ) . # $% $! %" 6 *%)!" (!&+= 3 5 , / % ,#!('%" $! " , , . . , 0 , B /, # , . 8 1 1 ' , 8 ( # ( , , 8 Jorge Ruiz Ramírez IM 5 , Tesis Doctoral , , 2 , 5 ( , 1 . #; , 3 H 2 5 A R( 2 , . 3 SU # - # + . , ( ( LH QG #- ( ( ) H8IG B 2 1 B8 ! IQQI"# ; 5 # , . , , ) , #; 5 , 5 , / ( ! 1' 3 # 0 , 5 / ( IQRG" 3 2 2, 5 U 3 ! , / U , , ( B / , ) . , , ,8 , 5 - IQQL"# , # , ) , . # ; ?C? ? #& &#4) *!$# )'! ! +"% $! ,%*,%" $! " ( , ) $% % * " 6! $%( " , , 3 ; B A 5( , 3 # - ) 1 5 1 8 8 1 . ( 1 #; F ! / K ) IQQL"# - ) ( A ' B 8 ( ) 3 1 , / , , . # Revisión Bibliográfica : ' Capítulo II , ) , 1 5 ) . , I $ , 5 FGGL"# , , 8 ( ; 3 5 , L 6+ JHa ! - IJ * F$ 9K# , 5 1 , , L B , Ja , . 1 ,B M . IGa ! 3 / IQQL"# "3 58 5 , , , 2 # , 8 , . ) , 5 3 B ) ) . , ) , # , , 5 - ( 5 $ ) 5 ) ) , ! , , / ( ) , ! , ,B 3 3 1 , : , #: + 8 , 8 3 FGGF"# , 5 ) ) ?A? ! %"% / %"'=" $% $ , "!)' = ) *#!)'% % !"' ,# #F &#4)1 . , 5 0 , B ) - , ) , 3 5 , 8 ! FGGL"# 5 H 6+ 1 , 8 ) 5 , 5 , , 5 5 ( ( , JHa , 3 ( . ) G QM B 1 8 5 / , Jorge Ruiz Ramírez IR Tesis Doctoral , !- # - , , 8 IQRM"# , !IQQQ" H , ) # 3 , # 0 3 , $ F 6+ , ) 5 $! 5 2! " , , ) , /5 , !] JHa" , ) , 5 %! 3 , / ) . 5 ) 8 , # - , , 5 ' # + ! 5 , 2 ( ( B , IH 6+ , , #; FGGL"# ; . 2 , , , ) 1 FR8LG 6+ 3 . , 2 , , 8 LK 6+ / , ( !C , : IQQR"# ( A ( , 8 2 , 5 ( # 0 8 , , B , 5 ( 5 , , , ( 8 IG ( 8 . # 5 3 8 3 , # ; IHa# 5 , / , ? ? !& $%=* $+( &#4) 1 - 5 , 1 , 8 , ) 8 3 / , , 8 , 5 , A # 8 IH 6+# ; ( 5 , ( 8 Revisión Bibliográfica ; Capítulo II IQRM\ $ , , ( 5 8 / 2 A. . 2 1 , , 8 IF 2, , 5 , # ; 3 2 8 3 IQQF\ + IQQL\ 1 ) 5' , 2 3 2 ( # : 1 # IQQK\ 2 C # IQQQ\ = FGGI\ + 3 # IQQR\ 2 IQQR\ IQQR\ / 5 , 8 L / , # 5 ) , , 8 IGGa # FGGH"# : 2 / # # IQQR\ C , 5 , , 8 , ! 8 B 3 , ( B , 5 , 3 , 5 1 # - , IQQL\ *'8 # IQQM"# - ( , $ IQ 2 ! IQQH"# : ( ( / B 5 # ; , B 5 , , ( 5 5 /, ( , A , 5 5 # , 5 / , ! IQQR"# ; , B , != 5 , 8 5 8 # /, , B ( , , :2 1 . !FGGL" 8 , 5 8 , , , 8 3 8 8 3 / 5 ) 5 ( ) 5 / 1 # !F Ia" , 8 Jorge Ruiz Ramírez FG ( Tesis Doctoral , . 3 3 ! 1 5 , . 5 8 ) 1 ' , ) # !IQRJ" 5 . , # IQRJ"# $ #"# , 5 ) ) / ! , ( / ( 8 5 8 5 ( 8 5 ) , , 3 B , 5 , , 5 #- ) 5 ) # . , ?G? )7 " $% ; ) ( ) ) IGGa 3 5 ,B , 2 , /, 2 ) 5 8 ( 8 ) ( ) 3 # - , 8 !:2 1 . FGGL"# : 8 B , 8 , ) , , ( ) ) , 8 ,2 ( S 8 ) , , 5 # - 8 , ) , 5 , IQRJ"# 0 ; 5 # M FJa ) 1 , L LHa , ' 1 8 RJ / , 5 , !IQRJ" 1 ( ) / ) , ) + # ) 2 , # 8 ! 3 5 ,B ) / ( ) 3 3 Revisión Bibliográfica , Capítulo II ,2 1 3 ,B 1 , 5 # # 1 - , , , 2 1' FI 2 6 # ) 4 3 5 / ( 8 5 ; 3 , , , ( 2 #* 5 A # / 8; # 5 8 !FGGK" 1 B MG 3 , . . , 8 3 S 1 8 5 , ( , 8 R ) 5 ! : ' :9 HKQF !IQQF" !IGGa *F 5 2 FGa +9F Z RGa *F"# ; , 1 5 B , 8 , . 5 )B , 1 ) 5 3 5 , 2 ) ) =2 ) , , , 8 2 , , . , , , 8 8 , 5 . , 1' 1 IQQQ"# ) / 5 ) . !FGGF" 2 5 # :. 5 8 5 , , # U 2 , 5 C? )B = ) , ( 3 , 8 8 / B 5 ) , % " - # ; , 8 / Jorge Ruiz Ramírez FF Tesis Doctoral / ,/ 1 , , , # - # , , 8 5 , 2 5/ 3 B , ! , "\ 2 )B , 3 / 3 3 B ) 8 3 5 , #0 1 \ ) " ; J 5 , , 8 5 / \ " ; , B B , " : , 3 , , ( , ( , 1 5 8 , # , , B # - 5 . 8 / , , . # ) ; , , 8 , , B 8 5 8 #+ . , /5 ) !9 1 , 1 ) , ( 1 , 3 , #7 ( , ( 8 , ( 8 , , # ; , 2 8 B8 , ;1 IQRR\ :2 IQQL\ + 3 1 . ' # IQQK" , 8 5 8 / 8 8 ) ) IQQG"# 0 , ! , / 1 , 3 , , IQRM\ + ,B , ' ! . IQRM\ $ # IQQK"# Revisión Bibliográfica - 5 Capítulo II , ( ) , 5 , 5 , % 1 ( 0 !07" 3 # - ' 2 , 8 , 3 , ) - P1 - / !- "c-P" 5 5 ' ) , ( 8 , ) 8 07 , ( # - 8 , !-P" ! -"c!CI - 8 , , ) # 5 8 * + S1 8 , ) , 8 ( 3 5 . 5 . ! # 1 # !IQQM" , . IQJR"# . ! " &'%(!" H+! C?B?B? '!(# 5 . , (#* +'# #F $ , , ) 5 8 # # !IQQK" . . ( 8 -c!- c-P" 3 # ) -Pc!07c-P" # 9 ) # 9 ) ) ( , ) . !IQQH" B 3 8 , + (#8!) 8!);'#&% ; ! -" , :!&' ) ( C?B?B?B? , - '!<'+( $! 6 *4) &+( $% !IQQH" 5 * ( C?B? 8 , ) 8 , 8 1 . ) . , FL # !IQQK" 1 8 !- c-P"# -Pc Jorge Ruiz Ramírez FK C?B?B? ? Tesis Doctoral ! !IQQR" %)'!)#$% 5 &%* %"#&#4) $! # !IQQQ" 1 8( " #)'( *+"&+ ( . . ; 1 3 5 , 8 ) ( ,$:=FK ^ M F , 3 ( 3 ,$:=FK ] H R# & .2 8 # 5 ! !IQRR" 1 ! , ) ) . ( /, 1 T" 8 # : 1 ' ( , 2 ,/ ,$:=FK ^ M F\ ' # =2 .8+ . 3 ( B ,$:=FK ] HR , !P " 3 2 8 M F# ; ! " 3 ,$:=FK ^ , 2 2 , 8 , # !IQRR"# 5 5 / , , , 3 ,$:=FK , , 8 # C?B?B?C? = ; = # !FGGL" . ,$ 5 8 # !IQQQ" 1 . , ( / , # ; 8 ,$:=FK ^H M , ) ] MG 3 , 3 d HM 5 ) # 8 3 ,$ d H M# ; / 5 , , ( 8 / 5 2 # # !FGGG" ) , / ,$ 8 # !IQQK" 5 , 8 . 2 ,$:=FK ) , / 2 Revisión Bibliográfica Capítulo II 8 , 52 # 5 5 5 !FGGK" / 8 # , 5 ( 8 . 3 " , , " 3 5 3 !RK Ka 3 5/ ! - ,$:= ^ 5 # :; ( ,$ . , , , 1 1 IQQR"# H HH !IH Ma"# + 5 5 2 , , , 8 3 ,$:= ] H HH ! . FH , , ) ! # IQQH" ) 2 8 8 # !IQQQ" 1 ) / # ) ( KH , , / C?B?B?A? %'!)&# ,$:=FK ^ M F 3 ( 3 8 ,$:=FK ] H R# * 0 # !IQQQ" 5 (%'!% 2'#&% , , 5 ( 1 8 , , / 3 ) , ( ,$:= !,$:=F ^ H R ( ,$:=FK ] M G" ) + 3 . , , , 5 2 IQQR\ ,$:=FK ^ M G# 9 /, C 2 C , , /8 ,$ ) 3 , , 8 , , 8 , , 1 2 , . # ( IQQR\ IQRK"# 7 , ( , 5 ! 2 2 ) 8 , 8 3 3 / ,$ ( ! ; , #4 IQQH "# , 3 / ) B Jorge Ruiz Ramírez FM 3 , / ) 3 Tesis Doctoral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evisión Bibliográfica 3 3 Capítulo II 5 , ) 1 A , 5 ) , 3 8 , 5 , 8 1 # 3 1 1 1 3 1 , . ! # IQQJ\ = 8 , ) 1 2 ) ( 3 , ) , 3 # + $ , C? ? ? (%&!"% '!&)% 48#&% ,B , 3 C? ? ?B? !* !( '+( 5 >+*!$ $ (!= ; - , , , ,B 8 5 8 , 8 2 5 , 5 ( 3 8 ) , !- , 5 3 , 5 #: 8 ! IQRM" 2 , , 8 , # IQQJ\ # FGGK"# ; 5 ( , , , ) ) ! # IQQK"# , 8 , ( 5 5 ) # '#7 - , 3 1 5 ) 2, , 8 8 # , ) B 8 3 , # IQQR"# 8 1 1 / 7 3 8 ) / , ) 1 5 , . ( 3 5 , ! !IQQQ" , ) FJ ) , , 8 , ) B , 5 8 Jorge Ruiz Ramírez FR ) . Tesis Doctoral , B / 3 5 , 2 , 5 ! , !:2 , # FGGL "# - # IQRQ"# * ( 5 ( , , / , 5 , ( 5 ) ) , $ . 1 3 8 , #; 8 3 - $ $ , JHa 3 , 5 + , 5 ) ) 5 5 8 ( $ , 5 ! 2 FGGI"# 1 #: JHa JHa 5 : # . , 5 , 8 #0 3 8 ) ( 5 ) 3 , ) ( ) # , , 5 5 JHa , 3 # FGGL "# - 1 2 , , 8 , ! FGGL"# 8 ! , 5 ( 8 , 1 , ) 2 5 , 5 , 3 ( 5 ) , ( , 5 ) ) 3 ) , $ , , 5 # : 3 8 , , 5 , 5 ) !9U # : ( , 1 3 , 5 ) # IQQF" $ RHa , Revisión Bibliográfica Capítulo II 5 5 5 , 5 3 ! " , 5 # , , , ( , , , JHa 5 3 # 5 ) RGa 3 / 2 # - 1 $ ) , , ( , 5 5 ) ) . , 5 ! " , 5 , , , 5 ( ! # - # FGGL "# 0 , , 8 8 2 8 , 5 ) , $ 5 , , 8 3 ) , 5 , 5 1 5 8 , 5 3 , / 5 ) , , 5 $ ) . , # ) , 2 , C?B? ? ? %)'!)#$% $! $ $ $! 8+ 5 " 8+ &'#7#= !0 :V , , , , 1 ) 28 2 '8 . ) # IQJJ\ :U(e IQQM\ # IQJJ = # IQQF" 2 FGGL"# = 1 3 , # FGGF" , . ! ! , ! , , ( 5 , , 3 / B B , IQQG" 5 I $#$ ' , 8 , 5 , , 0 FQ ! .8+ - # FGGK"# , Jorge Ruiz Ramírez LG Tesis Doctoral . # ( 1 ) ) !IQQH " : , # # = # !IQQH" 8 ) 5 $ 3 , !C IQRF"# ' , 3 /8 , ! . ( # IQRI"# ) . # !IQQQ" 1 ( , ) 1 . ( 3 8 ; ) C I 8 , 3 , ) 8 , 8 , 1 ( 8 3 .2 ! 1 !HJ G " FGGF"# - HL Fa"# + + .8+ , , 5 , 2 , $ # !FGGG" . ( B 8 5 5 8 ) ! IQ QR # FGGL"# ; KQ RIa #; K HF IK MMa# 2 - , , 8 25 # + 8 ) 5 5 3 , , , # 3 # . 2 8 , 2 , / , # !FGGG" , 3 , , B # - B , 3 , , 5 ) 3 /8 . 8 ) 5 ( 3 , , / 3 # , , Revisión Bibliográfica Capítulo II G C I# - I , 5 , + , B # !FGGF"# ( , ! , , # 2 !IQQF" A FGGG\ 3 ) , ) / , 3 5 B 3 ' 8 , / 8 , , IQRL"# , #, 8 * + ) 8 ) ) B ( . , , 5 # , , 2 2 5 !0 1 IQJF\ - V IQQI" , !: 5 ) 1, ( * + ( B 8 # - # FGGK : , 3 # IQQQ\ # FGGK"# , $ . 8 # / 8 / 8 5 , IQQI\ # IQQR ) . LI 8 ( # - 3 K Ha !IQRH" * + ( 8 Jorge Ruiz Ramírez LF 5 , 5 Tesis Doctoral 3 ( . ) , # IQQQ\ 3 8 Ma # !IQRK" !IQRQ" 3 # , , * + ( B , . SU A KR 5 MG LM SU 8 # ; , 8 * + 8 , ! 3 , " 5 5 ' , 8 , !$ Ha * + , #; * + / # !FGGG" 5 , ) 8 , , , ) , 1 , / ) ( / * + , ) +8 5 , ) , 18 / : , 5 , 5 - 8 / . ! ' V 5 . 5 B , # !FGGK" 3 8 . ' ) ( 5 # - , 8 B * + 5 M : ' 5 * + ) ! # IQQR\ / 8 La# # !IQQR" ( IQQI 1 FGGK"# 8 / # * + 8 ( 5 8 3 , 3 3 5 +8 , . 2 IQQH"# 5 2 # 8 * + , , / * + , 1 , # !FGGG" B , ) # FGGK" ) / 1 ( / 8 8 # 2, FGGG\ # . , 5 ) ' 8 8 Revisión Bibliográfica Capítulo II IFJ / # ( IR Qa * + " FK Ra - * + 1 ) ' 1 # IFJ , / , #, * + , ) 1 , ( ,$ ,$ 5 1 5 # IQQR"# , 5 * + ) ) ' * , , ' 3 ( ! 5/ LL 5 5 / G , 8 , 8 1 2 3 5 , 2 # - 8 8 ' # , , , ) . #- C?B? ?C? #!* % $! (%&!" $% 5 , 8 . 8 , ; 5 , , ( , #= , 1 ) ) # !IQQJ" / != ; 8 # IQQJ"# ' 8 . ) ,B , . / 8 ' 3 / , , 5 8 # ; ; '8 ) , !FHI . 5 , / "# - 3 8 #: 3 , G FHI ) , . 8 , ' 3 / , * + . 5 IQQG"# ( , 8 !9 8 , / Jorge Ruiz Ramírez LK Tesis Doctoral , , , 3 # !IQQK" 1 . B ( 8 ) 8 ( 5 , , ) # 1 5 5 FJL 3 # / IJQ / # * . # !IQQR" B , 8 # ( 5 C? ?B? )&%"'( $% , 2 !5 . . " 3 ' 5 , # & .2 , 3 , , 1 1 ! ( LRa , 5 5 5 8 , 2 , 5 3 , , , #; 3 , ! 8 IQQQ\ C , , / 8 # 1 , 5 B 5 FGGI" ! IQQI\ # IQQR\ IQQK\ # FGGG\ / 8 ; , 5 / 8 , , : ;, !; / !C FGGI"# # # IQQH \ # FGGK"# 5 . S $ # - % IQQR\ 2 , 5 ( , , !" (%, !* " $! '!<'+= , , 8 5 ) - ( 5 , ( !) 6 *4) &+( $% #0 , MGG / "# C? ? (#)&# 3 ( KFG / 1 2 , MGG KGa 5 , 8 KFG / ; 8 , , , 8 . " 5 , , 8 K#I#F !> : 8 Revisión Bibliográfica " Capítulo II , , ; : 8 3 8 , , + B, 1 5 / 3 ! " !C : C? ? ? !<'+( , )$ 5J % ; - , , % , 2, , 8 5 5 ) 1 , # , 3 5 ! # IQQH " ,$ ( ! / 8 , ( 5 ) 5 8 , ; , ! ( ) 5 8 2 5 , #; , , 2 ' 3 , 8 ) , FGGG"# 8 , # IQQJ"# ; ( * F$ 9K ! 58 , 1 , 5 2, , / 3 , 3 1 A , , # ; 3 , , " , 3 ! ) MF , # FGGK" , , 5 3 # IQQR" # IQQK \ , 58 ) ! , , , ,$ , 8 , S , 8 5 " 7 "'%" 8 IQQR"# ) ! # IQQR"# 8 " - , 5 B ,$ , IFa K#F#I 9 8 , , 1 ( ,B 1 LH A8 ) 3 3 .2 B ,$ ^ 2 2 , 8 8 ) , / # , , , 8 8 ' 5 8 Jorge Ruiz Ramírez LM Tesis Doctoral 1 5 , 3 , 2 , ! ! IQRF"# 8 C?C?B? ? 3 , , 8 "! 8 IQQI"# = , 2 , C?C? ;'%$%" +'# #F $%" $!'!(*#) &#4) $! ;'%$%" !* 2(#&%" 5 , # 0 ( '!<'+( ? , 2 1 , 8 , , 8 / C?C?B? #; ;'%$%" #)"'(+*!)' !" B , , - 8 8 !IQHR" 5 , 8 , 5 3 52 #- 8 : ) , 8 B8 . / , , , , ( C?C?B?B? : C?C?B?C? ;'%$%" :+)$ *!)' !" . , 8 ;'%$%" #*#' '#7%" 3 ) # 0 , , 2 5 ! 3 8 . , IQQM"# ; B " A C . 5 8 C IQMG 1 ) ! , . 2 ' , # , , 2 ( 8 #0 B 2 1 3 # , , 5 , , 8 . , % 8 Revisión Bibliográfica Capítulo II , , . : LJ C + 1 ) %b ; 0 ) = 1 ) 0 8 % L %0 . c+ 8 c; I b L 0 I C FS I I 1 0 b %0 . C b FS F# 25 , F , F 2 ) 8 5 C?C?B?A? # ;'%$%" H+2*#&%" 5 *#&(%"= &4 #&%" !:. . IQML"# ; U B 8 . , , 8 ) !C F"# , ! # ) 8 5 3 8 1 5 # . , ( , 2 8 , 5 ' , ) 8 ) ,$ , , 2 !IQQL" 3 / B B - V 3 / : K"# 2 B ) 5 , 2 , 8 8 / 0 , , /, 2 # - LR Jorge Ruiz Ramírez K# 2 , 2 / !:#)#&#4) "!)"%(# (9*!'(% +(!F Tesis Doctoral C , . 3 , !+ ) U IQJL" 5 ! !:#)#&#4) )"'(+*!)' )#$ $!" 8 :. . 8 : 0 " . 5 , . * V ! 1 IQQQ" C . 3 2 3 , , , ! IQJR" "'#&#$ $ ; 7 1 , , 1 IQQQ" ) ! , 8 ! 1 IQQQ" -F -I ; 3 , , 3 , f -FS-I ! $>!"#7#$ $ ; ( 3 IQJR" * , 8 , 5 !+ ) U IQJL" :. . 8 , 2 3 : , , , , 5 . ) 1 ( ) 1 5 1 1 %>!"#7#$ $ - 5 3 1 !:. . , ! 1 IQQQ"# c c #: ) 5 . ) # ; 2 ) . ) IQQQ" ! 8 . A U IQML" ! 1 IQQQ" B : ( ) ( #; ( 2 , ( ) f S ! B , , , ) IQJR" Revisión Bibliográfica #######+ Capítulo II LQ K# ( &'+( ,# #$ $ - 5 . :. . 3 5 # !+ ) U IQJL" : Fracturabilidad ; , 5 / ) , , . * V ! 5 , ! 1 ) IQJR"# . !:. . . c+ 1 ) c; ( U FGGF"# "'#& ,# #$ $ - / 1 %*%"#$ $ 3 3 , f0 , f 0 2 !+ ) , :. . U IQJL" - / 3 3 . c+ 1 ) B , !+ ) H# :. . U IQJL" B . ;'%$% H+# % 8@ F P# ?-; 8@ F ?-; 7*+ g* +9= ;: g* ( !:!(!)&# # !FGGF" 8@ F 8@ F P# 2 *: *: *: 8@ F 9*KLGI 9* KLGI 9* KLGI # !FGGG" # !IQQQ" # !IQQQ" IQJR" Jorge Ruiz Ramírez KG Tesis Doctoral , C?B?B?G? , # ; ;'%$%" , " $%" !) 7! %&#$ $ $! %" + '( "%)#$%" B , , 8 , , 2 8 # - # !FGGF" . ) 8 : C?B?B? ? # ; ;'%$%" :#"#% 48#&%" 8 3 ) : 3 ) 3 2 8 5 , 2 , 3 ) / ) / ! 8 5/ " , 8 ! 8 B , 8 B ) 5 8 , , , 8 , "# X , !IQRM" ) , 5 , ( - #= , , , # , B , 5 2 ( ) 5 3 ) 8 28 8 3 , 2 ( , ' 5 H# ) #; . C?B? ? , , 8 , . 3 , ) 5 B #- # , ) , )B 2 ;'%$%" "!)"%(# !" 2 8 8 . 3 8 #: 3 ) . 5 ' 3 , ( Revisión Bibliográfica M# 0 Capítulo II , . !"&(# '%(!" ( !:#)#&#4) C +(!F . , . # + +8%"#$ $ !:!(!)&# 3 5 ( : .8+ ( , , # !FGGF" # !FGGG" # !IQQR" # !IQQH" , 1 . # !FGGG" # !IQQR" # !IQQH" # !FGGK" A .8+ # "'%"#$ $ KI # : #,(%"#$ $ 5 , 1 3 1 , , / , 3 A # # !IQQR" # !IQQH" # !IQQQ" , # !IQQR" # !IQQH" 1 $>!"#7#$ $ , B # , ;5 %*%"#$ $ . 3 , , ) ) # !FGGK" # ) , !: B , , # 5 # !FGGG" # !"*!)+F ,# #$ $ , .8+ 2 8 # IQQL" , , , , , , / , , 1 . - 8 3 52 , 8 # , Jorge Ruiz Ramírez KF Tesis Doctoral , 1 , ( \ , , , 8 1 5 , # ; , , , 3 A 3 ) / #0 , B 2 , 1 , 8 , 8 5 5 . - 5 ( 5 B , /5 5 # B , /5 / 5 % 8 , 8 5 , , 8 / 8 # ; , ) B , ' 58 , 8 8 )B 8 . , , ) 1 IQQQ"# - 8 IQQK"# :. . U !IQML" / # / 5 , % 5 , - , , != ) . 0 8 , 5 ! # 8 , ) U 8 ) / IQQK"# ) 3 ) 8 ! , 5 , 3 , 8 8 3 , , ( 1 A ) # , # ' - 8 , 8 3 . ) 3 , ) 5 1 5 3 I# + 5 , # ) , 8 1 , / 5 . / . # 1 2 2 ) % , ) 8 % 8 8 Revisión Bibliográfica 1 ) # Capítulo II 2 % 5 / % A \, ( , 8 , 8 , ) 8 , ) 1 5 / , % , , ; , , M , ( - 2 3 # , 8 , 8 1' ) # , 8 1 # ;) 8 / # 5 , 3 8 ) 58 , , 3 8 B 1 5 1 , # ; ) HK B , , B IQQQ"# 8 , 3 8 . 8 1 FF # ! IQJR"# - C?A? ! &#4) !)'(! 1 8 5 2 S 3 ) 1 0 U !IQRQ" , ! 5 - B 8 ) . , , 8 B # L# 9 ( # / 8 / 5 5 5 1 ( , 3 , B , 2 / 5 # F# + 2 8 KL ,/ , 8 " *!$#$ " $! '!<'+( #)"'(+*!)' !" 5 "!)"%(# !" 2 + , ) , # 7 , , ( , # !IQQJ" Jorge Ruiz Ramírez KK Tesis Doctoral P G MI 5 IQRH\ . 5 . ( ( !- , # 9 h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evisión Bibliográfica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apítulo II 1 , HR MH8MR# # !FGGL "# , , # ; % ," 7 /# ; # + = # !FGGL "# ;55 5 , , 5 $ 5 ## # !IQQK"# 1 ,$ 1 5 # + , 8 5 # 8 . ( 8 4 ; / 8 ( ! + "# JR8RM# # + , # 5 1 ) 1 8 1 1 , 8 1 8 MH IFJH8IFRG# # -# 55 5 5 . , 5 V1 5 8 1 # * ' $ + , MM FJQ8FRF# # -# + # # !IQQH"# 1 1 1 55 . 5 :; 8 1 , #+ ' HF ML8MQ# # -# # !IQQJ"# ;55 5 , 8 1 1 5 5 8 1 5 1! * ' $ + , JK IQL8IQR# # -# :2 +# !IQQR"# 1 55 5 1 ,$ * + 1 ,8 ) 1 8 5 8 1 #* ' $ + , KM Jorge Ruiz Ramírez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esis Doctoral 3 ## 1 5 C 1 8 1 $ LJ FFQ8FKL# :# B # -# = # !IQQL" 8 1 ) , 5 , 5 C 1 8 1 # * ' $ + , ML MQ8JH# + +#\ 1 =# #\ # + # # !IQJF"# 1 ) . , , . , # ( , RF# + +# 1 =# # # :# !IQMQ"# 1 1 , 5 . , # , '" < ( FG LHH8LJG# + =# = # # . -# # # !IQQL"# : , , 8 1, 1 5 8 8 , 8 1 # * + $ HR QMR8QJF# + , # # C# / +# !FGGL"# ;55 5 5 1 ## MH MFL8MFQ# + # = / . -# 2 # B # !FGGH"# C 55 8 , 5 8 1 1 1 i j # # $ MQ JRQ8JQH# + , # # # 1 55 5 1 , , U 1 ! # $ HH FQI8FQH# + # # 3 # 8 # - , .8 +# / +# =# # !IQQK"# $ 5 5, , 5 1 # 1 $ Revisión Bibliográfica # $ LJ FIJ8FFJ# + ;# C # !IQRK"# : B , , 2 ( # ." , 57 ) 7 , FK LKJ8LHH# +) # # :. . U # :# !IQJL"# 8 , 5 , #* ) 6 $ K FGK8FFL# 0 1 -# X# !IQJF"# : 1 , 8 # , * = FH FLI8FKK# 0 # 0# :V =# # !IQQG"# $ 5 1 5 # * + % IF QH8IIM# 0 U # !IQRQ"# : S1 , , % , ) 6 $ FG #* I8FJ# ; ( # $ U # X # -# =# =∅ # !IQQQ"# + 55 5 , 8 ,1 , S 8 , 1 , Z, # * , $ JJ FKFR8FKLM# C 9# !IQRF"# 1 # + ,/ F# ; % ( / # ; ) B :# # ; , A LL8KK# C # !FGGI"# ; : % 5 ) # ." , / ( % 0 4 JH H8IG# C =# =# +# 2 C# !IQQR"# C U 1 5 , , , , # * , + ' KM KQRF8 Capítulo II KJ KQRM# C =# +#+# C# :, # =? !IQQJ"# + 5 ) , 5 , 1i j 8 1 5 5 V , # * , + ' KH FIJR8FIRM# C : # !IQQR"# , : # 9 FK * ) FGGL C : 8 : V % 1 ,%SS5 # S # , / 8; =# 0# 2 # !FGGK"# + , 5 5 ,1 , U ) , U 5 , 5 8 1 % 55 8 $ 3 ## HJ8ML# / 8 # & 8? 5 # , # - 3 + =# !IQQQ"# : , , 5 :, 1 8 1 5 5 ) 8 #+ ' MJ KFL8KFJ# / 8 # & 8? 5 # , # - 3 + =# !FGGG"# ) 5 1 , 0 $ - :, 1 8 1 5 5 ) # # $ HM I8M# / 8 # / 8 # & 8? 5 # , # - 3 + =# !FGGK"# 8 1, V ,$ 5 8 1 3 # # $ MJ MFH8MLF# =# # :2 +# !IQRQ"# ) :, 1 8 1 5 V 1 5 55 3 # >? ' ( #$) L JLK8JKG# KR Jorge Ruiz Ramírez ) ;# P# 0 ) ;# # # !IQJJ"# ;55 5 ) , V 5 U ) # * + $ KF ILFH8ILLG# .2 . # !IQQR"# , ( 1 #; %; % 8 / 2 # $ % &&' ( #$)# ; , # # # ; # ; #% # = 5 # =# III8IFK# # !IQQL"# + , / 8 5 ( . , # 0 # 7 ) 8 B # # -# # !IQQH"# : 55 1 1 5 8 $ KG FI8LI# 1 ## # + , # !FGGF"# 2 , , 8 ( #% IGH RH8QM# # + , # # !FGGF"# = ,$ 5 55 55 8 ) 1 H 6+# # $ MI FH8LI# # + , # # U V U ?# !FGGL"# 9 8 8 5 V ) 1 5 5 , # @' A $ - . # KMQ8KJM# # + , # # !FGGK"# = 55 ) 1 5 8 1 55 , ## $ MJ FGL8FGQ# =# -# !IQRM"# + ) 5 # ; % # Tesis Doctoral + #; # # # 1 # # LJ8IGF# -# # !IQQH"# 0 8 1 # +1 1 #+ ' ' JH KKQ8KHG# -# 0# !IQQQ"# ;) . % 8 8 ' 5 8 # ()%,# FG II8IL# -# # # 0# # !FGGG"# ;5 8 ( , , # (( $ ( # 8 * / # QK8QH# -# # & = # !IQQJ"# 5 ,$ 8 / ( # ()%, IR MJM8MJR# -# # # # !IQQM"# : 5 1 ,1 8 1 1 8 5 8 1 1 , , # * ' $ + , JG HFM8HLG# -# # !IQQQ"# 1 5 5 ,$ 1 , , 5 8 1 # # $ HF FMJ8FJL# -# =# 0# # !FGGK"# 2 , 2 # ; % 2 , # = / , 8 , A # ; # . # / # = # = # IQH8FIR# Revisión Bibliográfica $ # =# !IQRM"# P # ; % + 5 1 ; # # - V # 1 FJL8LIK# $ 2 . ;) # # # = 1 0# ; ) ,, : # * V 4 U# # !IQQL"# 8 ( # # 7 $%# Q IG8IQ# =# !IQQH"# 5 , 8 $ ;# $ , 2 5 ) $ Capítulo II # O 0 ( 8 # 7 8 # # =# =# =# !IQRJ"# ; ) ( # + ,/ # ;5 5 5 ( ) # 9 : ; %% 8 / # # =# $ # =# = 5 !; #"# # 8 # RJ# :9 !IQQF"# : % 8 ! : HKQF"# ) % 8 9 . 5 : 8 . # . # P X# !IQRH"# C % ) V# * ) 6 $ IM I8FR# 1 -# ;# !IQJR"# ) V 5 5 55 3 # 1 . '$ ) ' * I + 2# - V *# C +# P *# !IQQI"# 1 V 1 V , # 8 5 ) 8 , , # B '# * LGF RIQ8RFK# - V # # !IQRR"# +1 1 + ,/ ; # 5 K# ; % = KQ # # : #K # KL8JL# -# !IQRH"# k 1V 1 1 U # % =U 3 l ) 1V 1 1 U # 8 5k C 15 1 # X 1 QG MMGM8MMII# -# !IQRM"# k 1 1 U #+ ' ' MM KQM8HIG# - , # + # !IQQK"# = 8 1 , , 5 # # $ LM FGL8FLJ# - V =# # !IQQL"# , 5/ 8 , #; %. 7 # ; 6 / :# # ILH8IMJ # : :# # +# !FGGF"# ;) 5 , , 5 8 , ! " 8 1 3 #* + % 4 HL FJQ8FRH# = -# :2 1 . # =# : ( 2 # - , . *# !FGGL"# 5 5 V 5 3 5 1 # % (( # # * / +2 8; , A ! = # !IQQM"# ) , A , # = = IHM8IMR# = / A =# ;# =# +# + # # + ( # !IQQF"# = ,1 8 1 , 5 , 8 , 1 1 # + ' ' JF IHHL8IHHJ# = / -# 3 # + # =# # !IQQR"# ;55 5 , , , 5 8 HG Jorge Ruiz Ramírez 1 # # $ Tesis Doctoral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evisión Bibliográfica ( " ML KH8KQ# # # : 18 ) . ,, +# !IQQK"# 1, V 1 , ) , , 8 1 5 5 ) ## $ LR IIJ8IFF# # # !IQRG"# ; 3 8 3 5 5 m 5 8 ( # < 2 4I LH8KI# =# C . :# =# !IQQM"# , 5 8 V 1 , 5 8 # * ) 6 $ FJ HQJ8MFK# !IQMJ" 0 8 ' # FKRKSIQMJ# 5# L#IG# FI# ; + ;, A !IQMJ"# + ,/ @# + # =# $ $# '( # : # ?# !IQRQ"# : , U 1 8 ' # >D ( #$) + , 1 # JFH8JFQ# +# =# 2 C# !IQQR"# + , 5 , , . ) V 1 5 V1 , # * ' $ + JM IIJ8IFF# , 1 # !IQQQ" + ) 6 4 # 2 #; 8 , 1 *+# = 8 7: # .8+ =# # # + , # 0 #0# C 8 -# # # !IQQR"# 0 8 5 1 V 55 ) 55 , U # 2 &&'! ( #$) # .8+ =# # # C -# # # 0 # 0# !FGGK"# P 5 , U ,, *= # # $ 4 MJ IMQ8 Capítulo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orge Ruiz Ramírez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esis Doctoral ( V , ) , 8 8 5 1 # , '" E B 4 LQ JQL8RGH# 1 ) X# :U(e +# !IQQM"# P , 1 # * + % 4 FQ IL8FI# 2 C# !IQQF"# 1 . 5 8 5 , # ; * V 1 5 , # ; #% : C# # =# 2 C# C # =# ;++; =: *( # 2 C# !IQQR"# 8 , 5 ) ) , 5 8 , # # $ KQ IGI8IIG# 2 C# !FGGL"# = C %P 8 5 # + $ ) ' ( Q IJL8IJJ# 2 C# ; 1 0# # !IQRR"# ; 5 1 , 0 $ ) 8 1 ## $ I I8J# 2 C# C =# !IQQR"# 1 5 , , 5 ) ) , 1 , 5 8 1 # LR ." + $ LLI8LHF# 2 C# ;# C # !IQQL"# + 1 , 0 $ ) 1 , 5 8 1 # * ' $ + , MF IHJ8 IMI# +# :# !IQRI"# P ) 5 % 1 , # -# # U # C# : V # P ) % 5 5 3 # * V 4 U% # # +# : +# Revisión Bibliográfica # !IQQH "# ;55 , , . , , # ( " JG FI8LI# # # : 1 ) . ,, +# : +# =# !IQQH "# : 3 5 8 1 55 8 1 , ) , # * + $ MG IIRL8IIRM# Capítulo II HL # : 1 ) . ,, +# # : +# 0 =# !IQQR"# 5 1 , U 1 5 5 8 1 # * + B ' FF HL8ML# # P# !IQJH"# : 5 # * ) 6 $ M FHL8 FHJ# . 0 Materiales y Métodos Capítulo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orge Ruiz Ramírez HR ? Tesis Doctoral 5 B JGa *F" n : . , ( 8 ! I8 M" 8 L S F / SFK1 F SFK1 / B LG , ( % R % FG o" 8 IH 6+ LcI HcI H 8 K 6+SRGa $ \ , 8 !: +9- , , IG , ) , S FJ . SJG n % , ' K Ka# - FG IK , / / 5 3 # # !& $% )9#"#" $! '!<'+( - 5 , 3 1 - , B ,2 3 , , / , !F%H ,S," :, ,/, * c # ' ) , 5 ! 3 MG MRa ! " ! . , , 8 IQJR" , # . IQJJ"# - , , 8 , , / 5 ( . . !: ) ! 8 , = 7X" !C L" ) . ) 8@ F : 5 , . 8 , # 5 # . ( " / ) B 1 1 c , 2 IH 6+ , ! ' ) 5 IGcIGcIG IGcG HcI G IQRH"# - 2 8 / * !P 5 5 H ,/, ( HGa ! S# , 8 , , 8 Materiales y Métodos Capítulo III 5 " $F9SU , 28 % . ! " " 1 ! ) " 8U . * +" , 8 ! HQ * + 8 8 ' # ! "# )9#"#" :#"#&%H+2*#&%" "%,(! ! (!"'% $! *E"&+ % %)'!)#$% $! 8+ : ' :9 IKKF !IQQJ"# ; ,$ 5 C L# . , @ I# ,$8 o , ) . )9#"#" $! &'#7#$ $ $! 8+ / 1 5 &%)'!)#$% $! 8+ *+!"'( " +'# #F $ " / 1 "%,(! ( !IH SILH 1 8 - 8 "# " ) #F ( '!<'+( ; - . 3 , * ) !U SU , P ", 5 5 IGLWF p+ 1 , B 1 IQQM"# ! :9 IKKF%IQQJ"# B !U $F9SU 8U 8 ! :9 IRKI8I% , , IH ) ( 8U IHG * +" B !U 8 8 ; 1 . . 7 8 Jorge Ruiz Ramírez MG Tesis Doctoral ! 1 HG" IL#HGG , A LG ( RG8IGG 6+ 1 , + . . , , # C? - ( , IG : . 5 P1 * # HF# 5 , B , ) 5 IG - IG - , GI * A X*9L GI* 5 F , , R / , . LWF 6+ L / 5 , 8 2 8 JH8RHa # !$ " IG±I 6+ F / , 8 , $ ) ( " #)'( *+"&+ ( JG RGa IF±I 6+ , , : 1 , ! :9 IKKL IQJL" 3 1 / # ; 3 , ( !& $% , )8 5 ) / F , 3 # IH 5 1 , L±F 6+ , M 1 5 . , ) , , KH / # 3 2 1 8 1 ' B 8 2 , - U ( # ; * + 5 LG * *9F G F * + GF # 5B : ' 4 IG , 2 ( F , , , ( !=:8* + "# H H ( , A + , # . 0 , B , Materiales y Métodos Capítulo III C ) K K# ' . 6+ !-;" % - . # - -; #8 S , # 2 $FGS 1 . 5 ! S / 2 JHa $ "# - , ) , ) 8 QQ R W G Ia *F I 6+ , / / 5 IR / 3 8 , -*; ' 1 G I W G Ia 9F" % IKH FH 6+ M 5 ,2 F , . # 3 ! B ) , 5 ) ) L 1 ( . 1 8 5 R , , 5 1 -; 5 , ) G F8G L ) -*; HG±La $ K" IH±F 6+ / S# . ' , G F8 !-*;" R !C F±F $ ) GL * ; RG±La R / %- ; # . 5 MI ( 2 , / # ) .8 Jorge Ruiz Ramírez MF Tesis Doctoral # B ) 5 I S , 3 HGa , !$#$ , 2 8 $! " I# $! 5 $! %)&> " - / , , 5 , C H# I . , B F L K# 9 + !IQQG"# )9#"#" $! '!<'+( )9#"#" :#"#&%H+2*#&%" $! %" %*%" 2 1 5 ,/, , IGcIGcL 1 c " 1 , ! c 8 5 , . 5 3 / # 5 F±F 6+ , 8 , - 1 2 # : . % SH !:;= ; , A " !C H" %)'!)#$% $! 8+ , 8 8 2 5 , 2 2 1 # . , . , . . 2 B, ; 8 1 . # 5 = : , B 3 ) . 3 , / : 9 + !IQQG"# . B Materiales y Métodos Capítulo III Ka !G F ( ; , , 8 / * + . B B 1 IQQM" 8 3 ) . 5 - IGG - ) . , ) ) , . / M FH# 18 = 8 , , 2 2 ( " #)'( *+"&+ ( ! IQJJ"# : . / #'(48!)% '%' / 8 ! 9 + . B IQQG"# : , X( . A? M 1 C , ( 5' : k1 QRa !FG 5 -" , 2 . 5 5 , k1 ,$ , , ( 8 5 KFH"# ; 8 , , # ; ,$ 5 k 1 LIH" ,$8 , * 9$ LGa# ; !+ " ' ; , ) A ,$ 5 !0 1 / ,$ # - L 1 !0 LM , , KIG 6+ , A ( 5 # 5 8 , 8 FG 2 , I# 1 5 (%'!2) : , 1 $+ G FH *# , 8 5 ( GI - "# - ! :9 IRKI8I% 5 ML IGG - 2 FK 1 ) LM , 5 8 , IIWI Jorge Ruiz Ramírez MK Tesis Doctoral #"!I% !< !(#*!)' CG !()# !" BK !()# !" < !(#*!)'% C L !()# !" !) BK !()# !" < !(#*!)'% A L !()# !" Q R !) A> A> $;)'#&% &%) !" #!F $%" PG L !()# !" !)Q L !()# !" Q, !) A> Q R A >0 , . !()# !" Q R $;)'#&% &%) ,%( (6 *%)!" PG !()# !" Q R $#&#4) $! N N KN $#&#4) $! ! (%&!" (%) > "' CCOAN *!(* N N KN ! $!" #!F (%) * &!) $%" 7 &2% $+( )'! A $2 " (! ( &#4) $! *+!"'( " " (! !"'% $! *E"&+ % " !"'% $! *E"&+ % L'(%F%" AS S &* L'(%F%" AS S &* !& $%" > "' )#7! !" $!"! $%" !& $%" > "' )#7! !" $!"! $%" %)'!)#$% $! 8+ %)'!)#$% $! 8+ ( &'!(#F &#4) $! %" *E"&+ %" C ( &#4) $! *+!"'( " ( &'!(#F &#4) $! %" *E"&+ %" 5 = (%'!2) = = (%'!2) = 8( " =&%)'!)#$% $! 8+ 8( " =&%)'!)#$% $! 8+ M# ; 3 A , . , , 5 L K# Materiales y Métodos U IR Capítulo III ,$ d H J IR ,$ q Ra * +# , ' M F# - 5 , #7 ) , F . ,$ , ) ) ) / , !: +9- KH !; , FM / F $FGS . , ( 8 ) 8 FWF 6+ J# 8 , ) % . C K"# 8 L 6+S RHa $ " 2 FRQ / !; , S / ( , o, 1 L" 5 MF B , ' ) 8 ) , d HJ ,$ q , , ' ) , ( ; # 3 A C M# ; A , , ! ) 5 , 5 L " c F !,$" , ' ! C "# J# + 3 ' , A?B? 5 ( # C / , ' , , . 5 ( - . ' G GHa , 3 , - ' ' ( G GLa # 8 , 8 5 ) 5 X*9L 5 1 / ' , 5 A 8 ! , MH * *9F F H . H / KH / # ; , 1 1 , 3 8 Jorge Ruiz Ramírez MM . # 5 . 8 Q . !IR , , cF f LFK . ' cQ " 8 , + 5 , f / ) # A , L K ' T G UG ) ' ) L FJ L FJ L FJ L FJ L FJ L FJ ( !L ( Q ; ' # * f ' "# 5 ) , C 2 2 , , % ] I !: +9$FGS 3 / , F # o ; S / F±F 2 R# + # FL 6+SRHa $ " 6+ , R"# 8 ' K ) Q ' )'#$ $ $! I $#$ /N1 $ I S !C ; LGG" I R# 0 6+ HJ H±F Ha L±F G GI != , L , / , 5 F R# . . !& $% KGcFGcFG B ' Tesis Doctoral 5/ 1 , . ) FR M 8 . HM Ja# , , 8 Materiales y Métodos Capítulo III / 1 / . ) . . , , , 5( #7 . ) . ) ! " 8 , Q"# ;5 , , 2 , ,/, # ; F I 6+ , . !C ) / , , 3 5 3 IGcIGcIG MJ ) , , / LG / # ( , , 2 5 3 , F# )9#"#" $! '!<'+( )9#"#" $! &%)'!)#$% $! 8+ "%,(! 2 . ) " *+!"'( " $! *E"&+ % &+( $% +'#= #F $ " 5 2 I 6+ ( , 1 , : # - 8 5 / C ( ) #F ('!<'+( Q# , . , , ' 8 ) ,/8 5 9 + !IQQG" ! " ! , , !, , 2 # " I# ,/, " Jorge Ruiz Ramírez MR Tesis Doctoral . )9#"#" :#"#&%H+2*#&%" "%,(! ! (!"'% 7 8 HG" $! *E"&+ % &+( $% IL#HGG , LG ' 5 , . . 5 3 / B ( # !, , P1 , 8 , 5 " F±F 6+# FK 1 FK 1 %)'!)#$% $! 8+ - IGa# : ; , 8 IGG 2 ; ! 1 8 5 , , r HF ( HG : ' k1 , 9 + !IQQG"# B B . , * , ; ) ,$ ,$8 . - *# ! 2)$#&! $! (%'!% #"#" / 1 KGM8=L8:JSFH"# - . 8 ; * F8L B . / ,$ ! , 8 " I# 7 # 5 , #'(48!)% % (%'!#&% / ( ** 1 * ! IQMR\ + : B X , IGG - # IQJF\ + # FGGF"# !IQRK"% ( " #)'( *+"&+ ( IG ( # # IQQL\ : 1 ) . ,, ** . # $+ G I , !+ , 8 5 : RKFK" , F# - ) . !:1 8 ; 8 . 8 1 8 ; . 8 Materiales y Métodos Capítulo III / , A? ? MQ I# A (%&!" $% $! 6 *4) &+( $% 5 ( , 5 5 2 5 G GHa X*9L G GLa * *9F , F H Ra * +# , C , . , ) ) ) / , !: +9- FM F $FGS IG# # , o, S / 5 8 % ( ! !C , L F " JRWFa $ G GQ ) / , !C , 1 / 5 $ 1 3 , ( , , , , 8 ) H / # 8 # ; , 1 1 , JG8JHa B #- ( / IG"# ; , A LL S , , 5 ( FWF 6+ ) L J"# . 5/ 8 8 . ) , FL 6+S RHa $ " FWF 6+ , 5 5 ( 2 . ; 8 F 8 ) ! 8 , "# , ( , 8 3 3 ( ) , 8 , Jorge Ruiz Ramírez JG Tesis Doctoral ( , F 3 , 5 2 8 , 8 3 , ( # ; 8 , / $ 5 , , 8 ! Q# + Q"# , ( ' /$2 "1 /V 1 /N1 $% LL FWF 8 ! %"% MI FWF JRWF G ' C II# , 25 A ' ! C" ( !:=" !& $% B IL IGWF MJ HWF H !& $% IIG IHWF HJ HWF H !& $% C JF FRWF MH GWH G f f # !& $% ' Q , \ , \$ f1 !IR ( ) # . ; 5 , , ( 5 1 . , 5 0 , B , . 5 ( ' 8 II"# 5 ) / ' HR Ha# , ) 2 o, / !C ) ; LLWKa# ( / FF Ka FRQ / 5 c Q , ,B # - ' " , . KGcFGcFG c F f LFK 5 8 % ] I !: +9- 8 $FGS FL 6+SRHa $ " FWF 6+ 1 , 2 F S Materiales y Métodos 5 3 / 7 Capítulo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orge Ruiz Ramírez JF 2 1 , Tesis Doctoral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s # -# = 2 # 0 1 * 1 % = * (1 55 1 # MMI8 MJQ# . = ! W W ? D ; / 1 1 FKQ8FHK# , ! ) = : MQ Meat Science 69 (2005) 249-254 Capítulo IV JH !<'+(! ( *!'!(" %: $(5=&+(!$ > * " * !" $(#!$ ' $#::!(!)' !7! " " :+)&'#%) %: @ '!( &'#7#'5 )$ @ '!(&%)'!)' ? !(( + ? +#F= ( + , ) , V 1 + 1 F , , 5 V V 5 55 HGa IG V :, ! C" 1 # : ) 1 1 , # ) V " V V ) G#HH U ) , V ) 9P ) V FH 6+" 1, ) #C V 1 1, 1 $F9SU 1 V ! 8 V V1 1 1 ! 5 5 ) ) , V V V %+ "# + ! 5 V ! ! IJIFI = S , 8 *2(!F ? () + ? G#JG , ) , ) "# 1, V 1 V # P 0 8 1 B # Jorge Ruiz Ramírez JM Tesis Doctoral V B? )'(%$+&'#%) 1 , 5 , % 1 1 5 1 1 1 , ) # 1 5 1 , . V 5 1 1 1 3 1 1 8 ) 8 5 1 ) 5 8 1 1 : 1 ) . ,, FGGF" 1 V , 8 V1 8 !IQQH" V # 5 . # 1 MGa" 1 # s = # !IQQJ" ) 1, 8 5 8 C 5 1 C . 8 5 1 !HJ#Ka ) # HL#Fa\ 2 ] G#GH" 1 55 s 1 V1 1 1 1 1 ) V # 2 !IQQQ" 5 1 , V 1 U , 1 1 ,, 1 ! + 8 1 1 , 5 V ) V , 1 1 , 1 # # !IQQH" , P 1 ) 1 1 ) , # ) , = , 1 1, V 1 : ) V FGGI\ 1 ) 1 ! ) 8 " 1 1 5 5 ! # #% 8 # , IQQR"# ) s V ! , 3 ! $ V ) IQQM" . 5 1 s + # FGGL" IQQK\ : 1 ) . ,, 1 , ,, 8 V 8 , FGGL" ) 5 IQQH\ 3 FGGG\ 8 s : 1 ) . ,, 1 IQQR\ + s , ! ) 1 55 ! 5 !C , 5 ,, , 8 s 5 1 ( 1 1 # # 3 5 1 5 1 5 1 ! V" , . 1 1 1 FGGI"# 1 1 1 ) 1 , 1 V , 55 5 1 ) 8 5 5 5 !F#Ia ) Meat Science 69 (2005) 249-254 F#La" 1 V ) ! V , 1 ! FGa% I#IR U I#MQ U \ 2 ] G#GH"# * + 1, 5 5 V " 1 !1 ( 1, 5 5 8 V 5 !V IF8 1 8 1 V 1 1 5 1 ! ,, V ) V 1 , 1 , ! ,, V 1 # 1 1 # L c I#H c I#H IG , , 1" " LG V V , n ,, , 1 8 R ) , , 8 F \ "# 1 V , V 1 :#-# , U , S JG % K 6+SRGa $\ V % ) , , , "V 1 ,1 5 ) SFK1 : +9- !F c F , 1 1 S 1 # 1 !FG n L 1 Fa V 5 , 5 5 , , ) 55 C F 1 ) V 5 , 1 V 1 U 5 5 V " V # , 5 55 !$I # ) ! 8 C 1 , ) # !, , , 1"# V 1 5 ) 5 ! 1 5 , " V V ! C" IQRH" * 5 1 55 5 1 s 1 !$I $F\ $L $K\ $H $M"# 1 8 $F9 U , 1 $F" V 1 8 ) 1 , V 5 5 , ) # ! !$ 1 V 5* V 8 '!(# " )$ *!'>%$" 8 1 IQJJ"# 1 ) , 8 5 1 V V 1 HGG MG#MRa V V 1 , ) # IH±F 6+# 1 V 1 , IH 6+ 1 ? 1 8 " 5 1 , , V 11 5 5 1 8 JJ 1 !F%H VSV" !P 5 :, # 1 "# C V 5 1 1 5 s 1 !V 8 V .8 B +\ FG c FG c IG ) 1 + ) / !FGGG" 1 Capítulo IV F#G S + F SFK1\ N JGa 8 , 8 Jorge Ruiz Ramírez JR , Tesis Doctoral IH±F 6+ 5 U , 5 IK V 5 * + 1 V 1 5 V 5 , !$I 5 1 1 1 $F"# 5 1 , 1 ) ) 8 1 HGa 5 1 V 1 1 1 5 V V ) V 1 1 , , , V V V # 1 8, U 1 2 : , ; : " V V , 1 5 ) V 55 , F # +1 V " V 5 × 1 V% × ) # " ) - # 5 1 : 1 ) ) V * 8 $ HGG " 1 :V . 5 f U $F9 S U P ! - # V 8 5 IG c #C ! V" FH 6+ V 1 * ) 5l55 U IQJR"# 0 , IG c IG ! " V S P = , U )2, 5 ! 1 1 V ) #@ F !: , , ! S 1 1 5 # $ !>! G , V 1 1 5 # ! !) 6 # 5 F 8 1 ) " V G8L 6+ 5 1 5 5V , 1 , "# C 1 1 ) 1 ! f G 0 , , S# , V 1 V V 1 3 V 1 !:=: IQQF"% , !L8K , V 5 H @ # 5 , , 1 5 , ) # #C , V , V , , , 1 , V ) !IG V 1 1 , , , , 5 1 1" V "# + , # , U , 5 , 5 1 5 1 1 c 1 1" V $M V ) 5 * !3 1 $L $K $H 5 IG c H c IG 1 c V , 8 , !FH U " ! 1 1 V 1 3 8 1 , # 1 ) , 8 5 V V# 1 V " 5 1 ! , Meat Science 69 (2005) 249-254 Capítulo IV V 1 IGLWF 6+ V 1 1 1 1 ! :9 IKKF% IQQJ"# 5 1 , 1 1 =C" * + 5 f U $F9 S !U 11 hU ! 1 8 1 1 1 1 1 1 , ILH $F9" V 1 1 1 1 ) , 8 1 1 V 1 , K 1 # 5 IL!F" 5 8 1 : V ! U _ 1 5 1 U 8 U 1 1 5 1 5 # !: " U # C V 1 1 1 1 , ! + s IQQH"# ) 1 C 1 1 8 8 !:=" C? !"+ '" )$ $#"&+""#%) 5 8 :, 1 !0="# 5 : IHWF 1 , 8 t : IHa 5 , 1 ,1 5 5 5 t ! :9 IRKI8 :9 !IKKL% IQJL" I 1 V 1 5 5 V V ) I% IQQM" ! =C" * + IH#FKa !0="# : S V 1 1 V !IH M#QFa !;;+" * FGRFSQF"# V ,$ V 1 1 ! : " ! ! :9 IKKF% IQQJ"# $ #: L#RRa V 5 , ) ,$ 5 ) 5 1 5 V 5 , # = C , _1 1 1 1 IL#HMa V IQQI" # 1 1 C V ! 3 !0=" V1 !&! ' 8, U , ! =C" =C h U 1 5 ) 5 1 ) ,$ 1 * + "# 5 1 1 1 1 1 V1 1 1 V ) 5 $F 1 V ,$ ) " V , $I !V 18 5 1 V 5 !V 1 # $ JQ 1, ! V" 1 1 5 1 V V V ! " V 1 1 Jorge Ruiz Ramírez RG Tesis Doctoral I 1 1 1 5 , !0=" 5 # 8 1 1 5 1 1 !:. . 1 $ ,$ 5 !a 0=" * + !a 0=" $I M#RK M#QF IK#J $F M#JQ M#HQ IH#G $L H#QF M#JF IK#Q $K H#JJ M#JH IH#F $H M#IK K#MG IL#M $M H#QM L#RR IK#F 1 % 1 5 C5 C # I# 1 1 , V 1 C V , ) 5 , 5 V 1 G#HM U $F9SU V1 V ) 1 1 V V 1 V G#LH U $F9SU i1 V 1, 5 1 1 # 1 1 . IQRF\ C S F # f G#MK S f 8 # , 1 1 !2 ] G#GI" ) V 1 !MGa 1 ! C hG#JF\ := f hG#MG\ : C 1 1 1 V 5 1 5 1 8 , G#KJ ) G#MI !2 ^ G#GI"# V1 1 f G#JH !- IQRK" 1 , 1 f f hG#RI"# 1 : V 8 " 8 F 8 , FGGI"# 1 V KQ#J 1 V 1 1 ) 1 ) 1 , # 1 ) j1 V 1 , !C V 1 V ) , , 5 I#LG ) IH 1 1 1 5 !IQQH" 1 8 ) V 1 5 ) V V , V1 # $ V ) V 1 1 ) 1 1 5 , 1 G#JG V1 1 ) , # 1 G#QG 1 V 1 1 1 1 1 V 1 1 1 V C # I# 1 1 1 8 V ) 5 , V , V 1 5 # #% 1 U IQML"# 9 ) V , 1 V , 1 V 1, ) 1 1 V 8 , s: , #= V 1 1 1 # !IQQH" Meat Science 69 (2005) 249-254 , V ) 1, 1 , ) , 5 ) 8 ! V" 1 1 1 V 5 1 1 V 1 1 1 V 1 1 1 ! 1 , s !IQQR" s !IQQQ" V1 1 5 1 1 V 1 1 1 1 V1 1 1 , 5 1 V 1 55 1 ) , +) 5 ! 1 1 3 5 5 1 1 V V 1 ) " 1 5 5 5 s :. . ) \ U IQJL" V 1 1 , , 1 1 5 1 1 , # ) , , , V 1 1 1 1 , , 1 , 5 1 1 V 1 ) 1 8 V1 1 8 5 S , 5 , # 1 V1 1 5 # 5 1 8 ! " V" U 5 1 1 ,$ 1 ) 8 , ! 5 1 , ! V ,$ ) 1 5 C # F# P 1 , !IQQL" 1 1 V 1 5 #$ V ) 5 1 V ) 1 # ) V 1 5 1 1 1 1 1 ) V 1, V 1 $F 1 V 1 V , # 1 ) V $I 1 1 ) , # $ V ) ) V 1 1 V G#J 1 1 ) ,$ 1 1 8 C # F# 1,V 1 # #% 8 , 1 V 5 ) , ) 1 , V 1 1 :2 0 8 !C # I"# V V 1 1 1 1 V I" 1 ) V % $F V1 1 ,$ ) V 1 1 V V V 1 $I 1 1 V ! " V 1 1 RI # 1, # 9 ) 5 1 1 V V 1 Capítulo IV 1 1 V1 1 1 ) # 1 8 1 8 Jorge Ruiz Ramírez RF Tesis Doctoral HG HG $I $I $F $L KG $K $K $H $H $M $M LG $ LG $ $F $L KG FG FG IG IG G G#J G#R @ G V G#M G#Q G#L G#H G#J G#Q I#I I#L $I $I $F IF $F IF $L $L $K $K $H $H $M $M R +1 V +1 V R K K G V G#M G#J G#R F C # I# $ ! S ! U $F9SU $I8$M 1 # ) G#Q " C 1 5 1 # FGGL"# # !FGGL" 5 1 V1 5 1 V ! ) V ) ! 1 $F9S U 1 G#JF 0= 1 * + ) L#IH # V L#FMa 0=# 1 1 V 1 G#JG G#FJ U 1 V , 5 " V1 1 V 1 1 5 $ ^ RHa , ! " 1 1 1 U G#FK U , I#L 5 1 1 I#I ) IG , d G#JH# $ V ) 5 G#Q ") V HGa 8 V 1 F G#J F 55 55 G#H ! S , ) VV . G#L 1 V "# , @ G 8 5 8 1 55 5 55 Meat Science 69 (2005) 249-254 Capítulo IV G#MH %>!"#7!)!"" %>!"#7!)!"" G#MH G#HH $I $F $L $K $H $M G#KH G#J G#R G#L G#Q G#R G#J G#J (#)8#)!"" G#R G#M $I $F $L $K $H $M G#H G#J G#Q I#I I#L G#M $I $F $L $K $H $M G#K G#M G#J G#R C # F# + 1 ) U $F9SU 1 # , 5 1 1 , 1 1 8 1 V1 1 1 55 5 1 1 1 8 V # C V "# 9 ) 1 1 , G#HH U 1 1 V 1 8 , 8 1,V 1V V 1, V ! ) # A? &X)%@ !$8!*!)'" V 1 5 ) ) 1, , ! # $I8$M G#JG ) 1 8 I#L ) V 1 1 5 , V 1 I#I $F9SU V1 1 V 5 1 1 1 8 G#Q ! " IG ) HGa 5 , 1 G#J 8 5 ) G#H ) V , "# # G#L G#Q ) 55 ) G#H G#H G#K ) $I $F $L $K $H $M G#LH G#M (#)8#)!"" G#HH G#KH G#LH 5 RL 1 V :, 1 V 1= 5: 5 1 18 Jorge Ruiz Ramírez RK ! -FGGG8GHIJ8 K8GK , ( FGGGFGGGLLG C Tesis Doctoral * 1 . + ) / !C9* + " .8 / . : 5 # 1#0# C 7 ) U "# O V # ? !:!(!)&!" # !IQQI"# , ( , 1 # ( : , 7 ) C :, # # !IQQL"# # O / ( 7 $%#4 H4 L8Q# # !IQQR"# , , # # ; ( % / 2 # , ;, KK 1 + =: # % JI8RM# #s # !FGGI"# ;55 5 ) 1 1 5 1 , ## $ 4 HR MH8 MR# # !FGGL"# 1 ;55 5 # s + , 5 1 # ) 1 , 1 , ,, 5 8 1 5 # # $ MH IFJH8 IFRG# # -# + # s # !IQQH"# 1 1 1 55 . 5 :; 1 , # + ' 4 HF ML8MQ# # -# s :2 +# !IQQR"# 1 55 5 1 ,$ * + 1 , ) 1 5 8 1 #* $ + , JJ LRJ8LQF# =# +# !IQJR"# , 5 # + ) ' 4 > ? MF8MM JF# +) # # s :. . U # :# !IQJL"# , 5 , # * ) 6 $ 4 K FGK8FFL# + , # # s # !FGGG"# 1 55 5 1 , $ , U 1 # # HH FQI8FQH# C # !FGGI"# ; : % 5 ) # ." , / ( % 0# E A 4?D H8IG# : C . #+# !IQRF ! 1I% ' 9A : 5 ' ! 9 C %: # # !IQQR"# 0 2 ( #; ( % / 2 # : , ;, KK 1 + =: # % QG8IGM# , -# !IQJJ"# $ 5 , 5 3 #* . ' ' = B $ 2' ' 4 RI !I" RQ8 QM# -# # s # !IQQQ"# 1 5 5 ,$ 1 , , 5 8 1 # # $ HF FMJ8FJL# Meat Science 69 (2005) 249-254 :9 IKKF% IQQJ# 0 5 # ) % 8 9 . 5 : 8 . # :9 IKKL% IQJL# 0 5 5 # ) % 9 . 5 : . # :9 IRKI8I% IQQM# 0 5 1 8 I% 1 1 # ) % 9 8 . 5 : . # # :# 1 # # s : =# # !IQRK"# 0 , , 1 5 8 #* , + ' 4 > ILRF8ILRH# = # = ) # # = #C# + =# 0# s C# !IQQJ"# +1 1 8 1 ! $ " , 55 5 1 1 3 # # $ 4 KJ FQ8KJ# = # # + =# # s C# !IQQH"# + , 1 3 B 8 3 ,1 3 M , ( _; , # > ( $ # 2 ) # # !IQQM"# ) V% 1 ) 8 , , ) 8 1 1 % 1 , 5 1 # + $ ) ' ( 4 F MQ8JR# # # s : 1 ) . ,, +# !IQQK"# 1, V 1 , ) , 8 , 8 1 Capítulo IV 5 # $ .8+ # B RH 5 ) # 4 LR IIJ8IFF# # # + ) # s / +# !FGGG"# ,, 5 1 55 5 5 , # # $ LL QI8QH# :=: !IQQF"# $ # $ )!., A J I ! L#M"# : = : - : ; # :. . U # :# !IQML"# + 5 5 1 #* + $ 4 FR LRH8LRQ# # C =# C . :# =# s 2 C# !IQQQ"# 8 3 55 , , , U V 5 $ 4 HI 8 1 # # FHH8FMG# # # : 1 ) . ,, +# +# :# s =# !IQQH"# : 3 5 8 1 55 1 , ) , # * + $ 4 MG IIRL8IIRM# # s : 1 ) . ,, +# !FGGF"# = 5 $ 4 K LLI8LKL# ## P 5 P# :, P# ;# -# s # !IQRH"# : . 5 1 !+9: 8, ( QG QG "# 0# : s # -# = 2 !,, MMI8MJQ"# 0 1 * 1 8 %= * (1 55 1 # . Y W W = ; HIQ8HFH = 1 , = ) = : MQ / 1 Meat Science 69 (2005) 519-525 Capítulo V RQ (%:# !" %: @ '!(&%)'!)' )$ '!<'+(! #) &(+"'!$ $(5=&+(!$ %#) )$ #) )%)= &(+"'!$ $(5=&+(!$ %#) +#F= *2(!F ? !(( ? () + ? %+ ? #7 #+ + = 1 , # 1 , 5 V 1 5 5V ) 5 1 # ;8IJIFI ! " V ! " V 1 1 S 1 "# ) 1 U V V U , , = 5 8 , 1 , 1 # +5 !L8 1 1 1 U " 1 V 5 1 , 1 V 1 *+- 1 8 5 5 5 # "# ) 1 1 V ) 5 # :, 5 1 V 1 5 1 , 5 5V ) # V , 1 1 #$ 8 8 ! V 1 # 1 1 1 5 1 V # , 5 V % V # , 5 V 1,V 1 1 5 5 1 V # 5 +- V S 8 1 1 1 V K, " , V 5 ) , " 5 ! ! # V 1 , 1 !IGcIGcL V $ !+- $ !*+- ) 1 1 IH±F 6+ HG±La F±F 6+ RG±La X V # , 5 8 V V V 1 # :, , 1 "V 1 ( + ) 1 1 V # ) V 5 # Jorge Ruiz Ramírez QG Tesis Doctoral ! B? )'(%$+&'#%) 5 1 5 , 55 U s X 1 3 , , V .8 , s B s V 1 1 1 5 ! ! " ! FGGL"# 9 8 1 IQQK " , . ,, , 5 !: 1 ) 8 1 5 IQQH\ 5 : s C 1 5 1 1 ) , ; 1 5 !; : , "# 5 1 , 1 1 1 IFa# 8 , 5 1 1 81 ) 5 1 , 1 1 1 8 5 1 8 V # , 5 1 ) , :, 2 V 1 55 % 1 A s IQQQ" 1 ) , 5 # FGGF\ C . 1 1 1 s : 1 ) . ,, IQQK" + 1 , 1 1 ! 5 # U s 1 , V1 , : 1 1 # # FGGF\ : 1 ) . ,, , 8 s IQQR\ # V1 1 5 1 IQRH" 1 ,$ ! 8 / . 5 1 5 , # IQQQ\ :2 .8 , 5 s * FGGG\ , s IQQM" 1 5 8 1 V , IQQH\ !C / # IQQK \" ) IQQQ"# s .8+ + ) ! 8 . : 1 1 8 IQQR" 1 V FGGL\ 1 s / ! 55 V1 1 + ) s 5 55 1 , .8+ U IQMR\ :. . 8 IQML"# 1 IQQM" ! 1 # ,, ! 8 IQQJ" IQQI\ IQQR" 1 U FGGF\ = U V . s IQRJ\ :. . ! , 8 1 , !:. . s 1 1 1 !: 5 1 IQQR"# , 5 1 , 5 1 Meat Science 69 (2005) 519-525 , 8 Capítulo V , , 5 ) 5 V 1 , 1 ) 5 1 # 1 ( V ) 5 V 5 8 8 8 !5 A 8 8 , V 5) 5 + 5 , % IKH , JHa 5 ! ! V LG 5 * + X*9L # 6+ 5 G#F F , 5 V 55 F±I 6+# 1 8 U , 5 H 5 F V , $ 1 5 KH JG8RGa $# 5 V ) 5 IF±I 6+ 1 1 V ) , U , C 1 1 1 8 G#F8G#L 8 5 # S V 1 5 , F WF 6+ RGWLa $ 5 G#F8G#L S 5 5 1 U 1 8 " 1 R 5 *+-"# 1 , V 5 , U # G#Ia 9F 5 I 1 !5 V !L8 5 5 ) V 1 V $ 5 1 *+- V V L±F 6+ 5 U , 1 * + V 5 +- # 1 8 1 , 1 1 # V V ,, # , , 1 1 5 5 1 1 IG±I 6+ JH8RHa 8 , IHWF 6+ HGWLa L±F R 1 V U V 5 8 * *9F G#F 5 , ,, 5 5 S $" V 1 1 1 " V1 1 V V 1 F $FGS 5 1 !, , V % !+-" FH 6+ ! 8 !*+-"# V ,, '!(# " )$ *!'>%$" 1 1 1 R # 5 V ) 8 5 ) "# , 1 , 5 !I8 , ) V , ? K, QI M5 . V 1, 5 8 +- 5 , , 1 V ) QQ#R±G#Ia *F ,1 , 1 ! G#I ± 5 , : +9- 8 o Jorge Ruiz Ramírez QF , % F#M 6+ Tesis Doctoral F $FGS! 5 IR , , 5 5 IH±I 6+ 3 " RHa $" 5 FL u V 1 P8: ! ! ) 6 # :, = : SH :;= . " , 1 V , 5 1 "# , 5 8 V 1 5 1 ! 1 c V , , 1 c 1 , HGa 5 1 1 1 V ) 1 , 5I ! " 8 , 1 ) "# C 8 ,1 8 1 S# 1 5 " , 5 1 ) c , # !># 8 c # , , 5 1 = , ! X( V 8 1 V 1 1 ! 9 + IQQG"# 1 5 1 1 1 ! :9 IKKL IQJL" V 1 , , , V 1 IGLWF 6+ : V V : V ,$ V ,, 1 5 1 1 V 5 V " 1 , f U V IQJM" 1 V8 , ) 5"#" 1 " ! " 1 V ! ! V , 5 % 1 , , 1 1 ! fG V 1 ! 9 + IQQG"# P ?A >5"#&%&>!*#& 1 V , 1 $F9 S U 0="# V , 5 1 !0=" ! 1" , # 8 V 8 V V V 1 IGcIGcL 5 IGLWF 6+ # , , V V V 8 :V 8 ) # U , 5 8 5l55 U 8 8 8, U * 8 $ HGG V1 1 1 ! 8 - # " V 5 # 1 FH 6+ V 1 * ) # !7 ) 8 V V 1 1 , 1 , !+ !D$ , + , "# Meat Science 69 (2005) 519-525 V , 5 Capítulo V 1 , 1 , 1 V C + 9 5 1 : : , U V1 1 , # IQQQ"# , V 1 = ! -=" ! V *+-" # 5 55 8 0 1 1 1 ) 1 1 , * !FGGI"# 5 V , 8 1 5 1 I#KL W G#IQ !0=" 5 I#MIa !0=" , 8 V 1 $ 2 . * ) V , 5 , 2 + , 1 V , 1 1 5 1 1 4 1 V1 1 55 !IQQF" , 1 1 5 , , 1 #$ 8 ,1 1 , # 8 ) 5 # 1 V : s 8 8 1 I ) 5 !IQQQ"# C , , , s - , . !FGGG" U ! !IQJG" , , K#KQ W RG#IH W K#FIa 1 1 5 1 . 5 , #= !0=" * + II#FH W I#KLa !0="# 1 , 1 V 1 % ,$ M#GL W G#IF V s 1 1 5 V 8 $ 1 IQRM" ) 5 1 # 1 1 ) V , , ) 8 5 1 , 1 1 V 1 # 5 1 ,1 , , , S C? !"+ '" )$ $#"&+""#%) 1 , 5 ) #: 5 1 V 1 *- * 5 1 : :" 5 1 1 V8 1 1 ! 1 ) V , 5 ) U" 8 , , 1 # 8 V 1 1 ) V 1 1 5 1 : : , U !+- 1 - 1 # V 1 ) 1 !: : , QL ,1 5 Jorge Ruiz Ramírez QK Tesis Doctoral V 5 5 , , 5 1 ,1 ) 5 , 5 1 1 1 1 , 1 8 , 1 1 +- 1 ) 1 ) # 1 1 1 1 1 5 # 1 1 , , , 1 , , # 3 1 # :. . # # V # 1 U !FGGF" +- 1 , 1 1 V 1 $ V ) 5 # 1 5 "V 1 5 1 1 V1 1 1 *+- !FK#K 1 8 , 8 V 1 1 1 1 5 8 V 51 , ) 5 1 3 ) 1 5 8 +- 1 , 5 U V 1 5 1 U 1 ,, +- V V 1 ) 8 V +- V *+- 1 1 8 # $ V ) V1 1 V 1 1 V HGa 5 1 1 # 55 # : 5 , *+- 5 5 1 55 , 55 1 1 V V 5 1 , 1 V V *+- 1 5 5 1 5 ) 1 , , 8 8 1 , 5 V ) 1 1 ) , *+- 1 8 # $ ) # $ 5 1 # 1 V8 1 ) ,1 *+- 1 V 1 55 , 5 ,1 1 ) 1 V 1 , 1 1 1 ) 1 +- , V # 1 1 , 1 5 5 ,1 1 V1 1 ) 1 , 5 V 1 V V 1, 5 V1 1 V , #$ I 1 ) ) V # , 8 5 +- !RM#Q 1 "# Meat Science 69 (2005) 519-525 Capítulo V QH 1.2 1.0 Springiness 0.8 0.6 0.2 +F !IJa" Cohesiveness Chewiness 0.4 Hardness X 0.0 aw -0.2 -0.4 -0.6 -0.8 -1.0 -1.2 -1.2 -1.0 -0.8 -0.6 -0.4 -0.2 0.0 0.2 0.4 0.6 0.8 1.0 1.2 +I !JIa" C # I# V , 5 1 , ! "# 5 5 +- , , , : 1 f G#RQJ ! V1 ! K" : II#L 1 *+- V 1 V ) I#F 1 , 1 f G#RQL # $ V ) 5 V1 +- V 1 ) V V 1 1 # V G#RJG I#K fG#RJG\ ) 51 U , # 1 V 55 V G#R# 1 5 1 1 , , ) ) 1 , FR#K J#R 5 1 M#K 1 1 , $ 1, 1 1 ! f G#R" V1 1 # 1 V1 G#R# $ V ) 1 I#L f G#J 8 G#QIG 5 55 Q#H 8 V1 55 f G#R I#F FK#K ! " ) 1 V #C I#L 1 R#M $ V 1 V , 5 *+- V 1 +- V 1 V F 1 V 1, I" 1 V 1 1 C 55 8 # 1 FK#K , 55 , V # # V 1 1 V ") V 1 5 ! 5 1 V V 5 ! + , QM ± ) ($)!"" /" 1 #&! ) 5 %>!"#7!)!"" +- : *+- +- FK#K±M#I RM#Q±IR#J vv G#KH±G#GF G#LQ±G#GR Q#H±F#K FR#K±IH#J v G#HL±G#GK C M#K±I#H IQ#L±IF#F v A H#H±I#F II#L±H#I H#K±I#I H#F±G#I G *+- f 8 : If 5 : Mf v d G#GH# vv d G#GI# ! " V >!@#)!"" /" 1 ! " 5 1 <B +- : *+- +- : *+- +- : *+- +- : *: G#KH±G#GI G#HG±G#IG *: K#Q±I#K IJ#M±J#G vv RQ#J±I#F RI#I±I#F vv G#R±G#GL G#K±G#GK vv G#KR±G#GI *: G#HK±G#GL G#KR±G#GF *: F#M±G#K M#J±K#G v QI#I±G#J RM#G±G#H vv I#F±G#GH G#J±G#II vv G#HL±G#GK G#HF±G#GF *: G#HH±G#GF G#HF±G#GL *: I#R±G#L H#I±L#G v QF#F±G#L RR#L±G#H vv I#L±G#IH I#G±G#II vv v G#HH±G#GL G#HH±G#GL *: G#HM±G#GF G#HK±G#GI *: I#J±G#L L#L±I#L v QF#K±G#F RQ#L±G#J vv I#K±G#GR I#I±G#IH vv J#R±F#I v G#HL±G#GL G#HL±G#GM *: G#HK±G#GF G#HH±G#GI *: I#H±G#F F#F±G#L vv QF#R±G#F QG#L±G#J vv I#H±G#GQ I#F±G#IK vv R#M±K#L *: G#HG±G#GI G#HK±G#GL *: G#HG±G#GF G#HL±G#GI *: I#L±G#I F#K±I#I v QF#Q±G#F QG#R±G#J vv I#H±G#GM I#L±G#IJ vv \ +- f : ) *+- 8 # . V (#)8#)!"" *+- B , 5 8 \ f U $F9SU \: f Jorge Ruiz-Ramírez I# = 5 # Tesis Doctoral Meat Science 69 (2005) 519-525 , V RMa V1 1 5 ) , IQRM" 1 1 ) , ) 1 1 1 5 1 , 5 , 5 5 ) 5 1 5 V 5 ) # RG 1 1 , 8 1 5 : 5 5 5 KG G#L ! G#J" !C 1 1 5 8 V 1 , # 1 1 I#L P F" ) 1 G#R ! 5 1 # !FGGH" G # !FGGH" ) #$ V ) : MG ! " 1 -*+ FG V G#HH" -+ ) # 1 G#QG ) IGG 5 1 , P 1 ) V G#JG IFG 1 1 U ) ! " ( 1 ) 1 G G#MG " 1 I#GG FG 5 # G#RG # !FGGH" !U! " 5 : KG , 1 ) RG MG 5 1 V1 -+ -*+ $ !- 5 ) V !!U "" $ IGG $ P1 QJ IFG U V U FGGL"# $$ s Capítulo V C F#L !U $FGSU 0=" F# $ ! " , ) ; , ∆ !: 1 I#R ) V V ! "# f ! f 8 1 FGGH" , L6 L6 "# 5 , # 5 1 ! ) " 1 55 #% * + , , V 1 V IQJG"# 1# : 1 5 1 , 9 1 5 3 1 , 5 , IF 8, 1 ! 5 s 1 . .8+ # !FGGG" V 8 V 5 1, ! 1 5 Jorge Ruiz Ramírez V G#FH Tesis Doctoral G#QQ" 1 ) 8 1 U 1 # 5 1 U 5 ) 0.55 1, 0.50 1 1 1 1) 5 0.40 5 1 0.35 # >! ! # ' 1 0.45 + 1 !U " 5 1 5 -+ -*+ 0.60 8 1 1 0.65 , G#H8 , V ) QR ! 5 V V 0.85 8 5 1 V ! " 0.30 0.80 ) 0.90 P 1 1 % f 0.95 ) G#MH -+ -*+ G#MG L6 ) L6 ! + 1 V 5 G#HH 1 % 6# L 1 V 1 1, V G#HG ) 5 ) F 5 G#KH + 1 !U " C G#KG V 1 1 1 ) # G#LH 5 1 , G#LG 1 1 V 5 1 #- 5 ! , V F 1 55 8 1 ) 8 5 8 " 5 V ! 8 1 V "# : V , , 5 5 1 .8 / . ! , 1 5 + , # 8 1 U V 5 1 ! "# , 55 , 5 ) s 1 \ 1 , V #9 ) V 5 ! 1 , 5 1 , 5 FGGL " V1 1 1 5 # !FGGL"# 1 L# + 1 ) ) ) ! " V ; , , f ∆ , 1 ) ! f L6"# B 1 I#R C 5 8 $ G#R I#L !U $FGSU 0=" P F# 1 V 5 G#L ! " 1 1 5 ! e s Meat Science 69 (2005) 519-525 Capítulo V *%$! # $ %&'() *+) *+,- :%(> ($)!"" )$ &>!@#)!"" ,! ? '>!* '#& &%>!"#7!)!"" : L#IHHI 8R#GQKF H#IQQL G#RQ J#FQ G#GGI G#GGMF 8G#GFRL G#GRQJ G#QM K#KI G#GGI FM#HRI 8MK#QHJ LQ#JMQI G#JR F#IL G#GGI 8G#GRRR G#FRIQ G#GQLM G#QG I#KH G#GGI G#LQKK I#GGKK 8 G#LL G#GK G#GGI G#LRGM 8 G#KM G#GK G#GGI \ V ! 5 1 U $F9SU # ) IQQF\ # s 1 5 5 # ) # V 1 1 V 1 1 1 1 V V !L8 C 1 1 U" 1 5 ) / !C9* + " 5 V # 8 8 1 V V V # 9 + !IQQG"# , , ' # !IQQI"# 8 E # IH 1 ; ,, QLI8QLH# , ( , # ) # &X)%@ !$8!*!)'" 5 : !:!(!)&!" 1 5 5 1 V C .8 O 1 1 ) #$ V 1 1 * + U 1, V 1 1 FGGGFGGGLLG 7 ) . # :, # -FGGG8GHIJ8 K8GK / . 1#0# + 1 V "# . A? %)& +"#%)" 5 5 : ! , ( 5 , , 5 1 = 1 V != 1 IQQH" :, e # FGGL " 1 G#IIIM "# # 5 IQRR\ : 1 )$ # $ () +:%( %$! @#'> %$! @#'> ($)!"" /. 1 >!@#)!"" /" 1 %>!"#7!)!"" 4V ) 4 , F f 55 =:; f QQ 1 # !IQQJ"# ;55 1 5 1 5 ' $ , 7 ) C ; , A# -# s 5 , 1 5 8 1#* + # IGG Jorge Ruiz Ramírez , JK IQL8IQR# # -# s :2 +# !IQQR"# 1 55 5 1 ,$ * + 1 , ) 1 1 5 8 1 # * ' $ + , JJ LRJ8LQF# A :# + ;# +# =# 0# s =# 0# !FGGF"# , , U 8 # &-' ( #$)# # IFM8IFJ# *# +# !IQJM"# 5 1 ' , #? ) ' ( # 0 * V 4 U# 1 :# B # -# # +# s = # !IQQK "# ;55 5 ,$ , 1 5 5 1 8 1 #* H !L" FHJ8FJG# # + 1 :# B # -# # + =# 8 # +# s = # !IQQK "# V , 3 55 1 8 1 # # $ LJ FFQ8FKL# C # s * # !IQRH"# + /, ( , 2, # ." , 57 ) 7 , 4 FH !I"% IIJ8IFK# C : # !IQQR"# , : # =# # # s - , . *# !FGGG"# P 55 ' 8 # ( Tesis Doctoral A V (U 1 * # LFG# # !IQQH"# : $ 1 * # * 8 # , -# s # 55 1 1 # $ KG FI8LI# # + , # s # !FGGL "# * + , 8 55 55 8 8 ) 1 M 5, U1 ## $ ML FQ8LK# # + , # # s U V U ?# !FGGL "# 9 8 5 V ) 1 5 5 , # @' A $ - . ,, KMQ8KJM# -# # # s # !IQQM"# : 5 1 ,1 8 1 1 5 8 1 1 , , #* ' $ + , JG HFM8HLG# -# # # s # !IQQQ"# 1 5 5 ,$ 1 , , 5 8 1 ## HF FMJ8FJL# $ $ 2 . # * ) # s 2 C# !IQQQ"# - , ) 1 V :, 1 5 , % 8 , U 8 ! # $ HI IFL8IFR# $ # s* 0# !FGGI"# ) V 5 1 ) 1 5 5 1 # * ) 6 $ LF FIQ8FKF# :9 IKKL% IQJL# 0 5 5 # ) % 8 9 . 5 : 8 . # # Meat Science 69 (2005) 519-525 - -# !IQRM"# k 1 1 U # + ' ' # MM !K" KQM8HIG# = 1 # : =# = 1 4# $# X V :# s V =# !IQQH"# * 8 ) 5 !* " 1 # * * $ + $ ) ' KF !M" KLM8KKI# = U V . $# s # !IQRJ"# + ) , 8 . 55 # $# # = U V . !; #" C # * V 4 U% = 0 UU # LLH # # s : 1) 8 . ,, +# !IQQK" 1, V 1 , ) , , 1 5 5 ) $ LR IIJ8IFF# ## # !IQQM"# ) V# 1 8 ) , , ) 8 1 1 % 1 , 5 1 # + $ ) ' ( F MQ8JR# *# !IQRM"# C 1 # + $ ! !+ ,# IG ,, FKM8LGF"# * V 4 U% 1 ) 1 + , # +# s . -# !IQJG"# +1 ) , , 8 , 5, U 8 U , # * + $ KH FKR8FKQ# e P# s : # !IQRR"# + 5 1 , , # + ' ' MR !II" IKFL8 IKFM# .8+ # # + ) # B # s / +# Capítulo V !FGGG"# ,, 55 1 5 + . QI8QH# .8+ # !IQQR"# : IGI 5 5 , # '( # =# 4 LL s 1 1 % 5 , + . '( HL8HR# .8 / . # : @# # !FGGL"# ;5 ) # + ) / +# 5 5 # # LI # s ( / # # 9 (( * / # +2 8; , A # : 1 ) . ,, +# 0 =# * + -# # .. -# s # !FGGF"# 5 V , , 1 # # $ 4 MG JJ8RL# : @# .8 / . # # s # !FGGH"# , 5 8 1 # , 55 ) 5 5 V ) V # # $ MQ FKQ8FHK# : 0# s 4 4# !IQQF"# 1 3 5 1 5 , # $# # : 1V . s # P# $ # 2' ' !+ ,# IM ,, MMQ8MRQ"# * V 4 U% = 0 UU # # : : # !IQQQ"# : 8 : -!@ ! + *+% : : # : # s e P# !IQQF"# 1 5 5 V ) 8 # + ' ' JF !II" IGF Jorge Ruiz Ramírez IHKJ8IHKQ# :. . U # s X 0# !IQML"# + V 5 1 5 # + ) ' IJ!I" JK8JR# :. . U # !IQMR ! + V ( ) #+ ) ' 4 FF QRI8QRL QRH8QRM# :. . U # !FGGF"# , , # + 3 2 IL FIH8FFH# #C =# C . :# s Tesis Doctoral C# !IQQQ"# 2 55 , , U V 8 1 # # $ FHH8FMG# # # : 1 ) . ,, : +# s =# !IQQH"# 3 5 8 1 55 8 1 , ) , # * $ MG !M"% IIRL8IIRM# 2 3 , 5 HI +# : + . W W = D ; / 1 1 HJQ8HRJ , ) = : JG Meat Science 70 (2005) 579-587 Capítulo VI IGH ! '#%)"># ,!'@!!) @ '!(&%)'!)' &%)'!)' #) $(5=&+(!$ *+"& !" +#F= *2(!F ? () + )$ '!<'+(! ? !(( ? ! ; #+ S IJIFI = 1 5 1 , 1, V 1 V 8 "# 7 1 # 1 , 1 1 #$ 5 5 1 1 1 V ( + 55 ) 1 ,$]H#J , 55 ,1 5 ) 1 V 5 # $F9SU V 1 5 1 V " V1 5 1 * + , 1 G#M 1 1 5 1 5 1 1 ) , V 1 , V V ,$ * + V * + 1 5 8 5 , # , 5 1 1 1 V V 1 ,$:= ^ M#F# V ,$:=# %$ ,, 3 # 1 , # V , I#L 1 1 ) KH HQ#Ja V ] G#M 1 1 1 V U # 1 G#R 1 1 , U V 1 1 1 ,$:=# 0 8 ) FR#H 5 1 V V ,$:= 1 1 # 5 1 , V , 1 F 1 5 1 V 1 ,$^M#F FWF 6+ 5 5 1 !U 1 V 5 1 , FK81 , U # ,$ IR 1 V 1 , 5 * + U 1 ) %+ + :, V ( *!'!(" 1 IGM Jorge Ruiz Ramírez Tesis Doctoral :2 B? )'(%$+&'#%) 1 8 1 ) , 1 1 1 5 1 , 1 5 V1 1 , , , # 5 1 1 V 1 = 1 , s C 5 / . : 1 5 ,$ 1 5 5 1 ) , 1 V 1 ,$FK1 ] M#G 8 1 V 1 V 1 ,$FK1 ^ M#F 1 1 V 1 ,$FK1 ] H#R 55 .8 9 , V 1 1 1 55 1 ) 1 V 1 # ,$ 8 V1 1 IQQI\ ! # IQQR\ & 8? 5 , / 8 8 s- 3 IQQQ\ s !IQQF"\ 8 8 55 + # KH # ) 8 1 ) 5 5 , * + * + 8 # 5 5* + 1 C . V 1 ,$FK1 ^ M#G# FGGH" 55 8 V 1 ,$ ] H#M# 9 1 1 IQQH\ 1 ,# ( 1 !IQQQ" 5 FGGL\ : s , ) 1 1 .8 / . , 1 ! s 55 1 ,$FK M#G V 1 1 ,$F1 ^ H#R + 1 H#M 1, IQQH\ : ( 2 8 s 1 5 !C : 1 ) . ,, # P1 2 !IQQQ" V 1 :2 1 . 1 , , = 55 , 1 ,$FK ^ M#F 1 . 5 1 ,$FK ] H#R 1 1 55 3 FGGI"# " 1 1 5 1 ! - , . !FGGL" , 1 V 1 1 1 = 5 5 1 1 1 # # 1 , V1 1 1 8 1 1 1 5 , , 5 1 !IQQQ" ) 8 , 1 !IQQR" / 8 FGGG\ & / 8 8? 5 Meat Science 70 (2005) 579-587 , s - 3 + FGGK" 1 ) , !:2 s+ Capítulo VI IGJ 5 IQRQ"# # !IQQR" 5 55 1 ) V !:=" FK81 , ,$:= , 1 # 1 1 ,$:= 1 ,1 ,$ 8 1 ,$:= ^ M#F# V 7 # $ V ) V 1 1 1 5 MG = ) SU KR SU # 1 V1 5 * + Ma 1 ) FGGK"# V La ! + ) , 1 s 1 1 , 55 V 55 # 5 * + 1 5 * + ,$ V 1 V 1 FG HG RG # ) 8 5 !: +98 % F#M FL 6+SRHa $" ! , , "# 5 1 , !K F F 5 "# , 1 5 1 !: +9- 8 o , % ] I $FGS FL 6+SRHa $ " 1 , V U 8 ) , U A 1 1 V '!(# " )$ *!'>%$" , 8 FWF 6+ 5 KH # F $FGS V . ,, 5 )8 1 o , 1 U , U , S 5 V !*fLFK" V ) 1 5* + , 1 1 V 1 , , 5 1 1 5 5 X*9L G#L * *9F - V 1, ) V 1 . 5V HGJ := , 1 5 !+ 1 B 1 55 ? "# 5 G#H 8 1 :# ! C" LM " ,$8 1 5 55 , :, * + V KGM 5 :V . + # !FGGG" 5 V 1 ! IR#Qa ) FK#Ra 5 * + V1 1 ,$:= ] H#J , 5 ! #$ ,$ = 5 IR 1 FWF 6+ 5 5 1 ,1 1 F S IGR Jorge Ruiz Ramírez # ; 1 , V 1 ,, LWF 6+ HJ#HWF#Ha 1 ) , $ ) 5 FR#H V , V 5 ,1 , U 1 i F±F 6+ 5 3 1 # :, SH :;= ! i1 ) , V 1 1 5 1 IQJR"# , V , fG"# V 1 5 V V V 1 , 5 V ,1 8 V 5 1 , 1 8 V 1 !0=" !@ f U $F9 S U 0="# # ! : V 1 V 1 1 IQJM"# 1 5 1 1 1 S# , V F#K 1 1 C 5 IGLWF6+ 1 HGa 5 1 1 ! , V , , 1 ,, ! 9 + IQQG"# P ! 1" V 5 I 1 1 , V 1 IG c IG c IG 1 , 8 ) , , 1 5 V 5 1 # ) 1 8 V V 5 , 8 5 1c1 1 j! 1 # 1 cV 1 " 8, U U , 5 , 1 !># IQJR"# * , , , 8 " V V ) 5 1 5 1 5 = : 5 , 1 , j 1 !7 ) ) 1 1 # ! ) 6 1 5 V 1 . i1 , 5 LG , ) V 5 j# + 1 8 U , o" "# , U 5 , !: +9- " ! 5 , 1 ! ! HQ#Ja G#I±G#Ia 9F % 1 V 1 # 1 ) QQ#R±G#Ia *F , V , V I S V , V ! " 1 $ Tesis Doctoral = ! *" Meat Science 70 (2005) 579-587 X( 1 1 #$ 5 ! =C" 1 :9 !IQJL" V # V Capítulo VI IGQ 1 ) 8 , 3 / . V 1 !+ " 9 + 1 5 , !IQQG"# P 8 , V #* 8, V V 1 !X ) 1 X( 1 , 1 # ) ) 1 V V 1 1 % 1 I 8 = ! -=" , * + 55 V f IT 1 " , # 1 V 1 ) 1 * + ) 5 0 1 1 , 55 KH 1 1 8 1 5 , 1 1 1 ! # # V # , C? !"+ '" )$ $#"&+""#%) IQQQ"# 1 ,$:= , : : !: : - " , 1 # !I" 1 5 !D$ 1 5 : :"# ! f IT 1 ) =C 1 ) # 6 !0=8* + "# , 5 1 , , U V 5 , !* c M#FH" V 1 1 1 V 5 1 * + V 1 1 8 IQRK" 5 1 5 P1 5 , V 1 1, I + + ⋅ 6 + ⋅ 6F = V 5 ! ; , V , 1 *- * , : :# V .8 : !FGGH" , 8 !;3# I" , , , !0=8* + "# 1 ,$ V , 5 V ,1 ,$:= , ) 5 # V V 1 1 1 ,$:= V ,$:= 8 , , 1 5 IIG Jorge Ruiz Ramírez V Tesis Doctoral =C V 1 , V * + 1 1 V !$ 1 * + V 1 55 55 V 3 1 V 1 V ,$:=# 1 55 5 1 := U U 1 * + V1 * + 1 " 5 5 * + 5 ! ! / . 8 1 # 1 # V V , C , 1 V ,$:= , := 5 3 =C 5 1 # I! " 1 V 1 5 1 1 V 1 5 = 1 1 , C 1 ! ^G#GH" 1 V 1 55 # 1 ) 1 5 1 1 =C !4 FGGH" !: 1 C V G#HM , 55 1 # FGGH" 1 , 1 , 1 1 8 55 5 1 1 1 1 V 1 55 .8 V ) , ) =C # ] G#GH"# ] G#M 1 # #% G#R F"# # 1 , s * + ! 51 55 1 V !,$:= 5 8 1 # FGGF" C , G#R , V1 1 =C = 1 V 1 V # 8 V 1 FGGF\ = 5 1 1 1 # ? 8 1 V 3 U8 8 #; , 5 55 8 * + !+ 1, I#L 1 1 # 1 V 1 5 , , 1 1 1 1 1 I! " 1 V V R IQRG"# C V =C 1 1 5* + 5 V 1 1 1 ,$ , , ) !C # 1 ) 1 1 5 V 1 # * 1 ) 5 1 1 , , ) IQMG"# ) 1 55 1 1 # 1 5 1 # !IQQJ" V1 1 C 1 1 , 8(%+ %* %)!)'" N !" '!$ @ '!( &%)'!)' (%'!#) / = 1 / = 1 B Z / J 1 :#) / 1 = V 1 55 0= f 0 0=8* + f 0 5 8 ? $$!$ !7! ? +"& ! #8>/P G? 1 %@ /Q ?R1 %@ / N1 !$#+* / N1 JH#GR ±G#MG JL#LH ±G#QH JK#HQ ±G#QF JK#KM ±G#RH JL#MG ±I#KR JK#GR ±I#FQ JK#LH ±I#GJ RM#MM ±F#LJ RK#RL ±F#JG RH#FH ±I#JQ RH#RG ±F#JQ RM#IR ±L#LK RK#KG ±F#LM RJ#GQ ±F#FQ R#KQ ±I#JH IL#RF ±I#HI M#LH ±G#IK IH#KG ±H#HR IG#LM ±F#IG IL#GG ±G#QQ H#RI ±G#GM IK#LL ±H#JF Q#LK ±I#HJ IL#MR ± I#FQ M#IF ±G#LH J#RQ ±G#JM IG#GK ±F#GG IL#LI ±I#HF M#GR ±G#FR IM#IG ±G#QJ R#QG ±F#MR IL#FH ±I#FL M#GH ±G#FJ FG#MG ±I#FJ IG#LR ±F#GJ IL#MJ ±I#LI M#GM ±G#FQ IK#MM ±H#HI R#KR ±I#JJ IL#IM ±I#LL M#IG ±G#LG IH#GJ ±H#HI 1 V !V 1 1) " 5 55 #8>/KN1 ? Meat Science 70 (2005) 579-587 = I !±:#0" 5 ,1 ! ] G#GH"# # # Capítulo VI III IIF Jorge Ruiz Ramírez F !±:#0" 5 * + = Tesis Doctoral =C +(!$ *+"& ! !a" 8(%+ / V, U C I! " 1 $ 1 - V IH#FK ±H#HR IK#QG ±H#JM J#KQ ±I#GQ Q#KJ ±I#RG 1 5 , , 1 ,$:= , I#L 1 1 , ,$:= ,$ , , !$ IQRM" #C , ) 5 IQQM" 1 1 s ) 5 F! " F! ""# 1 * + 1 , 8 1 5 1 * + 8 , 5 !:1 X V1 1 1 1 5 1 1 s 1 , 8 5 P 1 1 1 , # 1 IQRJ" IQRM" 1 1 1 * + V 1 1 1 1 V # IQQR"# $ 1 ) V U !X V ,$:= ! !C , 1 V 1 1 ,$ 1 ) 8 5 ) !4 1 1 V ) 5 5 I#J# 1 1 U 1 1 1 1 ,$:= 1 5 ) ,$ 1 V 1 # 5 1 1 1 G#M V G#M V V ! !IQQQ" V1 5 1 1 1 55 55 1 V ,$:= 5 V1 , ) 5 ) 1 # !IQQR" 1 1 ! ] G#GH"# V 1 1 1 1 55 , , "# 5 1 Q#HG ±I#JH II#FH ±F#GJ !,$:= 5 = IH#HJ ±H#HJ IL#JH ±H#MI ,$ , / V 1 1 1 ,$# 1 V 1 := " 1 # 1 , $ 1 - V = V 1 55 1 $ 1 f ,$:= ^ M#F\ - V f ,$:= ] H#J# 0= f 0 # 0=8* + f 0 # := !,$:= ) 55 5* + ! U s ) IQQQ\ FGGG"# Meat Science 70 (2005) 579-587 Capítulo VI IIL IFG IGG .8 : 0 8 / . # !FGGH" # !FGGH" ) ! " RG $ MG KG FG G G#LG G#HG G#JG G#QG ! " KH KH KG KG LH LH LG LG FH FH FG FG IH IH C IG I#LG I#HG I#JG ! " !U $F9 S U 0=" P $ I#IG ,$:= ] H#J ,$:= ^ M#F IG := H H G G#LH G#HH G#JH G#QH I#IH G G#LH I#LH G#HH G#JH G#QH I#IH ! " P C ! !U $FGSU 0=" I# $ .8 / . , 5 ) # ) V # FGGH" 1 ) V ! " P 8 # ; , 1 N I#LH !U $FGSU 0=" , f 8 !: # FGGH" Z N ! " , 1 ! " ,$:= ! " 8 8 IIK Jorge Ruiz Ramirez Tesis Doctoral KH KH ,$:= ^ M#F $ ! " KG ,$:= ] H#J KG LH LH LG LG FH FH FG FG FG#QHa * + IH IH#KIa * + IH IM#JQa * + IG FG#FMa * + J#LIa * + IG R#KJa * + H H G G#LH G G#LH G#HH G#JH G#QH I#IH I#LH G#HH G#JH G#QH I#IH I#LH ! " P C F# 1 , *" 1 IGGu!* * S 1, V * + 1 , V 1 # : U !IQQR" 1 5 1 * + V1 V # 1 * + 55 , 1 55 5 1 V V 1 1 5 , !C # F! ""# * + ,$:= ] H#J ! "# , 1 1 !Ra" 1 V 1 ) 1 V * + 55 1 V 1 Ha 1 5 ^ G#R ] 1 !IQQJ" # !IQQR" V1 1 1 * + 1 1 ,$:= 5 55 5 1 * + V 1 55 % IM#Ha ) IQ#Ha IR#Qa ) FK#Ra V ,$:= !C # F! "" 1 Fa V G#H# 1 , 1 1 1 V 1 ! ]G#GH" 5 ,$:=# V 1 5 ) 1 V ) 1 55 1 1 1 * + B 8 V ,$:= : # !FGGG" # !FGGK" !U $FGSU 0=" 1 1 * + 55 , 1 1 P 5 ) ) V 1 ,$:= ^ M#F ! " 8 1 ! " !U $FGSU 0=" 1 , ) 1 # 1 ) 1, ) V 1 1 Meat Science 70 (2005) 579-587 5 1 * + 1 V ) 1 5 V1 1, V 1 ,$:= 1 1 1 55 1 V 1 5 1 ,$# 8 1 1 , 5 " 5 1 3 !I" =:; 5 * + ,# C 1 1 V , ) I#L ! ] G#GH"# V Ha 55 1 5 , 1 8 ) 8 V 1 8 1 5 1 , 1 , 5 i1 5 5 5 5 i1 j U 1 G#MJ G#MJ 1 U !IQML" V1 1 8 5 ) # 1 , ) 1 V1 U # 0 8 5 , V V , ) , ,$:= :. . 8 H! " 1 V ) 1 , 55 V 1 1 1 ,$:= , ! " 8 1 1 V ,$:= 1 5 5 ,# 5 1 V ,, # L! " ) 1 1 Fa * + 1, ,$:= V , V ,$:= 5 1 V 1 , 1 8 1 1 1 ,$:= 5 * + ) L# 1 V 8 H! " , " # V , , ! 1 5 1 1 5 ) 1 := L! " , 1 V 1 ! ^G#GH"# ! =:;" ,$:= 1 55 5 , C 1 , 1 8 8 1 1 1 V 1 , 1 C 5 5 * ) FGGL"# 55 ) 1 s 1 H! " , 1 ) 51 1 5 5 , 1 L! " , ! 1 C # 1 1 , + # 8 ! !FGGH" / . 8 1 1 ) 5 .8 , 1 V ) # !FGGH" 1 ) 1 * + F 8 V # 1 V 1 V G#M : 1 1 * + 1 , ) ) V G#M 1 ! Capítulo VI IIH 5 5 ) j# IIM Jorge Ruiz Ramirez Tesis Doctoral L ; ) 5, 5 1 8 51 V N ! N ! " G#GILH 8G#GHRM G#FLFG G#RL L#MI C := G#GGJK G#GIRR 8G#GLFH 8G#GRLM G#FGFG G#FMFQ G#RG G#RM L#QK L#FM ,$:=]H#J ,$:=^M#F G#GGII G#GFQI 8G#GGFI 8G#IFQL G#IMHK G#LIKF G#RI G#RM L#QG L#IJ 5 &%)'!)' / Q ?R1 Fa 8G#GGFM 8G#GFFI G#GFHF 8G#GIHR G#GRLK 8G#GKQF G#FKFK G#GQMK G#IKRG G#RL G#RM G#JQ F#IR L#LI K#FR 5 &%)'!)' P G? 1 / Fa G#IQHL G#GFJG G#GGFL 8G#MRLM 8G#IKJJ 8G#GGHH G#JJQL G#LHFK G#IJKG G#JQ G#QI G#RM L#FF F#LJ L#HH !)!( *%$! 5 *+"& ![ 5 4 Ha Ra Ha Ra , # ) v Cf 5 1 F \ 55 \ := f 1 , ,$:= 5 1 , 5 1 , V1 5 1 1 1 :. . V ) V , U 8 1 , ,$:= 1 5 , 1 ) 1 1 !IQQQ" 8 1 5 1 V 5 1 1 !IQJH" # !IQQQ" V V 1 ,$F1 ^ H#R 8 ,$FK1 ^ M#G# * ) i 1 # , 1 V 1 1 , 1 5 V 1 V 1 55 ,$FK1 ] M#G V 1 , ) 8 ) 1 5 V 1 1 1 ,$:=# 1 5 , ,$FK1 ^ M#F# $ V ) 1 !IQML" 1 1 1 , KH V 1 ,$FK1 ] H#R 1 IQRM" V1 1 1 \ =:; V 5 U ) 5 # , !$ 1 ) 1 1 FS j I 1 5 8 Meat Science 70 (2005) 579-587 Capítulo VI IIJ G#MG G#HH G#HG G#KH ) G#LH + 1 G#KG G#LG G#FH G#FG G#IH G#IG G#LH G#HH G#JH P G#QH G#MG G#HH G#HH G#KH G#KH G#KG G#KG ) G#LH + 1 := G#LH G#LG G#LG G#FH G#FH G#FG G#FG G#IH G#IH G#HH G#JH !" G#HG C G#IG G#LH I#LH !U $F9 S U 0=" G#MG G#HG I#IH G#QH I#IH I#LH ,$:= ] H#J ,$:= ^ M#F G#IG G#LH G#HH G#JH G#QH I#IH ! " P C L# ; , ! " , ! " !" !U $FGSU 0=" 1 ) ,$:= ! " P !U $FGSU 0=" 5 1 ) ) ) 5 8 I#LH # ) V V IIR Jorge Ruiz Ramirez G#MG Tesis Doctoral G#MG ,$:= ^ M#F G#HH G#HG G#HG FG#QHa * + G#KH G#KG ) IM#JQa * + G#LH G#LH G#LG G#LG R#KJa * + G#FH G#FH G#FG G#FG G#IH G#IH G#IG G#LH FG#FMa * + G#KH + 1 G#KG ,$:= ] H#J G#HH G#HH G#JH G#QH I#IH I#LH IH#KIa * + J#LIa * + G#IG G#LH G#HH G#JH G#QH I#IH I#LH !" P C * + K# 1 V 1 U , I 1 5 ) 5 ) 5 , , , V 1 * + !C K! " K! " 1 ) 1 5 1 8 ! 1 5 F " 5 1 8 , * + ,$:= V 1 , ,$:= 55 1 ! =:;" 1 V F K# 1 1 ,$:=# 55 # 1 1 M! ""# $ V ) 1 1 8 5, ) V 1 1 1 1 , M! " # , 5 1 1 ,$:=# 1 1 1 1 ,$ 1 1 , ) 5 1 1 , ) V 1 1 ,1 ) V ,$:= ] H#J ! "# , 5 1 ) !U $FGSU 0=" 1 V 1 , 1 # 1 V , ) P 5 ) V 1 ,$:= ^ M#F ! " ) : 1 FS ) 8 ) ! " !U $FGSU 0=" 5 1 , Fa 5 * + V 5 V 1 1 1 5 V 1 # %)& +"#%)" * + 1 * + 1 ) ,# 1 1 , 5 := Meat Science 70 (2005) 579-587 Capítulo VI IIQ G#JG G#MG G#HG :, G#KG G#LG G#FG G#IG G#LH G#HH G#JH G#QH I#IH I#LH !" !U $F9 S U 0=" :, P G#JG G#JG G#MH G#MH G#MG G#MG G#HH G#HH G#HG G#HG G#KH G#KH G#KG G#KG G#LH G#LG := G#FH G#FH G#FG G#FG G#IH G#IH G#IG G#LH G#HH ,$:= ^ M#F G#LH C G#LG ,$:= ] H#J G#JH G#QH I#IH G#IG I#LH G#LH G#HH G#JH G#QH I#IH ! " P C H# ; , ! " , , ! " ,$:= ! " !" !U $FGSU 0=" P 5 8 ) # I#LH !U $FGSU 0=" 5 , ) ) V V IFG Jorge Ruiz Ramirez Tesis Doctoral G#JG G#JG ,$:= ^ M#F G#MH G#MG G#MG FG#QHa * + G#HH G#HG FG#FMa * + G#HH G#HG IM#JQa * + G#KH IH#KIa * + G#KH G#KG G#KG G#LH :, ,$:= ] H#J G#MH G#LH R#KJa * + G#LG G#FH G#FH G#FG G#FG G#IH G#IH G#IG G#LH J#LIa * + G#LG G#IG G#HH G#JH G#QH I#IH I#LH G#LH G#HH G#JH G#QH I#IH I#LH ! " P C * + M# , C V 55 * + V V 1 1 G#M 1 ) 1 5 * + 1 1 55 5 5 5 # &X)%@ !$8!*!)'" P :, V 1 1 = U V 5 : 8 . % FGGG8FGGGLLG" 5 .8 / ._ 7 ) # !:!(!)&!" V 5 8 5 O 1 , ) C9* + 1 1 1 1 ,$:= ) 5 ( 1 * 1 V ,$:=# C 1 85 V V 5 5: !, 1 V 1 ,$:= ^ M#F# 5 C ) , ) V ,$:= ] H#J ! "# -FGGL8GKMIF * + 1 , !U $FGSU 0=" 1 1 # 0 8 V 1 1 V P 5 ) V 1 ,$:= ^ M#F ! " 8 ,$:= ! " !U $FGSU 0=" 1 B 1 ) # + ) # # # s . # !FGGK"# : 1 5 1 % 5 5 , # # $ MR KH8HI# 9 + !IQQG"# , E , ' # IH 1 ; ,, QLI8QLH# # !IQQI"# , Meat Science 70 (2005) 579-587 Capítulo VI IFI K ; ! ) N " 5, V 5 1 5 1 ) , ! " %>!"#7!)!"" G#IFJQ G#FRJR G#MI G#GHRK C := G#IFKK G#ILIG G#FRKG G#FQFL G#MG G#MF G#GHJQ G#GHRH ,$:= ,$:=]H#J ,$:=^M#F G#IGRM G#IMFL G#LLLF G#FFIK G#JH G#KM G#GHGR G#GHMJ * + !,$:=] H#J" Fa Ha Ra G#GHKR G#GRQI G#GQRG G#LHQM G#LMRG G#LJRM G#RH G#JM G#JM G#GKLI G#GHIJ G#GKIJ * + !,$:= ^ M#F" Fa Ha G#GKJK G#IQQG G#IMIH G#LGKI G#IJHK G#FJGR G#MK G#HH G#MH G#GHLG G#GLMH G#GKHM G#IKQH G#LFKM G#MG G#GMJI C := G#IFIF G#IJMH G#LHLR G#FQMR G#MH G#HH G#GMHI G#GMRL ,$]H#J ,$^M#F G#IGKK G#FIQF G#KGHG G#FIGM G#RH G#LL G#GKKQ G#GJGM Fa Ha Ra G#GKQI G#IIGQ G#IIHQ G#KKLH G#KIIG G#LQQQ G#QG G#RM G#JJ G#GKFR G#GKFH G#GKLI Fa Ha Ra G#GRKR G#FJFM G#IRQF G#FRQR G#IKGR G#LFMK G#JF G#KL G#MH G#GKFQ G#GLJI G#GHHG v Ra (#)8#)!"" v ,$:= * + !,$:=] H#J" * + !,$:= ^ M#F" , v Cf 5 1 \ := f \ , , F 55 5 ( # # !FGGL"# 1 1 ;55 5 \ =:; ) # # 7 ) C :, # # s + , 5 1 8 O 8 8 # ) 1 , ,, 5 # # IFRG# # !IQQJ"# ;55 1 1 5 $ 1 MH IFJH8 -# s , 5 1 , 8 5 8 # 5 1 IFF Jorge Ruiz Ramirez 5 1# * ' $ + , JK IQL8IQR# # -# = ( ;# s # !IQQF"# ;55 5 ,$ 1 5 8 1 # >-' ( #$) FFQ8 FLF# # -# s :2 +# !IQQR"# 1 55 5 1 ,$ * + 1 , ) 1 5 8 1 # * ' $ + , JJ LRJ8LQF# *# +# !IQJM"# 5 1 , # ? ') ' ( #0 * V 4 U# =# +# !IQJR"# , 5 ) ' 4 LF MF8 # + MM JF# P# =# IQJH# ,, 5 , 5 8 5 ) #* 6 $ M HL8RF# + U8 U =# = # s ? # !FGGF"# U 3 , 1 8 5 8 1 5 0 # * , $ RG QRR8QQM# C # !FGGI"# ; : % 5 ) # ." , / ( % 0# E A 4?D H8IG# C # # !IQRG"# 0 55 5 1 5 , U# * + $ KH IJKG8 IJKK# / 8 # & 8? 5 # , #s- 3 + =# Tesis Doctoral !IQQQ"# : , , 5 :, 1 8 1 5 5 ) # + ' MJ KFL8KFJ# / 8 # / 8 # & 8? 5 # , # s - 3 + =# !FGGK"# 1, V ,$ 5 8 1 3 # # $ MJ MFH8MLF# U # $# s X 1 =# !IQQQ"# ;55 5 , 5 5 , w8 , # * , $ JJ FMRH8FMQF# U # $# s X 1 =# !FGGG"# 18 ) 5 w8 , # * , JR FLLM8FLKL# $ -# # # 0# s # !FGGG"# ;5 8 ( 8 , , # (( $ ( * / 8 QK8QH# -# # s # !IQQQ"# 1 5 5 ,$ 1 , , 5 8 1 # # $ HF FMJ8FJL# $ # IQMG# 1 5 1 #," + . '4 IG LHH8KML# $ # !IQRM"# C , , 5 1 5 1 # # + ! !+ ,# K ,, ILH8IQQ"# ; # 1 # 7 ) 5 7 # ?# : :# s : U $# !IQQR"# ;55 5 V 1 * + 1 5 Meat Science 70 (2005) 579-587 5 1 Capítulo VI IFL # 1 #+ ' $ 4 MK !M" QMQ8QJF# :9 !IQJL"# :98IKKL% 0 5 5 # ) % 9 . 5 : . # X # !IQRK"# # ' # +1 1 % ; $ V - # X 1 =# !IQQM"# 1 5 1 1 . , 5 # # $ KL :IQL8:FGI# = U # # # :# + :# *# = X 1 C# X# ;# # s : ) # P# !FGGF"# + 1 ,, 1 3 55 1 , U 1 1 , # * RG QQJ8IGGK# , $ = -# :2 1 . # =# : ( 2 # s - , . *# !FGGL"# 5 5 V 5 3 5 1 # (( # # * / +2 8; , A # = # # + =# # s C# !IQQH"# + , 1 3 B 8 3 ,1 3 M , ( 8 _; , # > ( $ # 2 ) 8 # = # = ) # # = # + =# 0# s C# !IQQJ"# +1 1 8 1 ! 1 " , 55 5 1 1 18 3 ## $ KJ FQ8KJ# .8 / . # : @# #s # !FGGL"# ;5 ) ( # / 9 (( * / # +2 8; , A # / . # : @# #s # !FGGH"# 5 5 V V ) 8 8 8 ## $ 4 MQ HIQ8HFH! :2 +# =# # = 5 # s + # !IQRQ"# ;55 5 ,1 ,1 1 ) 5, , ! # $ 4 D4 & &?! : : # !IQQQ"# : : -!@ ! + *+% : : # : @# .8 / . # #s # !FGGH"# , ! 5 8 1 , 55 ) 5 5V ) V ## $ MQ FKQ8FHK# :1 # P ?# # s ) # !IQRJ"# : , , 5 ) , !1 , ,1 1 " 55 * + + +F #+ # M IQQ8FGJ# :. . U # :# !IQML"# + 5 5 1 #* + $ 4 FR LRH8LRQ# # C =# C . :# =# s 2 C# !IQQQ"# 2 3 55 , , , U V 5 8 1 # # $ 4 HI FHH8FMG# # # : 1 ) . ,, +# +# :# s =# !IQQH"# : 3 5 8 1 55 1 , ) , # * + $ 4 MG IIRL8IIRM# 4 -# # s - 4# # !IQRM"# ;55 .8 IFK Jorge Ruiz Ramirez 5 ,$ Tesis Doctoral , ) #* HI JJK8JRG# + $ . 0 A W W ! !/0 = 1 ? A . D ! !/0 ; 1 # # 1 , , , = : # Meat Science accepted Capítulo VII IFJ ::!&' %: &%)'!)' )$ (%'!% 5"#" #)$!<%) '>! (! '#%)"># ,!'@!!) A @ '!(&%)'!)' )$ '!<'+(! ( *!'!(" #) )$ *+"& !" #) $(5=&+(!$ > *? ? +#F= *2(!F ? () + ? !(( ! ; 5 1 , + :, V 1 1, %+ "# 7 #+ S IJIFI = 1 ? 1 V ( + 55 5 ,$ V U 1 1 IR 1 ,$]H#J 1 FK81 51 ,$^M#F 1 5 1 V 5 FF#K , 55 HR#Ha# 1 V 5 1 , V 1 , V1 5 1 #: " 1 1 V 1 1 1 ,$:=FK * + 1 , 1 , , 5 !1 1 5 , ) , , 1 , F 1 # %1 #0 8 1 #$ 5 5 1 RG ) 1 !IGG× V ) , V V " 5 1 V , V1 1 1 , 8 1 8 1 V , 51 V 1 # V ,$ * + 8 # 1 , 1, VV U , 1 V 1 5* + , 5 , ) 1 , 5 1 , V 55 1 1 5V 1 1 , # 1 # , V ,$:=FK 1 V S 1 5 ) 5 1 , , FG HG , V * + 1 !,$:=FK"# 1 1 # ) , 8 V , , 8 IFR Jorge Ruiz Ramírez Tesis Doctoral 1 B? )'(%$+&'#%) , 5 1 :, i 55 8 .8 IQQR"# , ! 1 5 1 IQQR\ / 8 & 1 1 1 , s - 3 + FGGI" ) 8 IQQK\ : 1 ) . ,, IQQH" ! IQQI\ :2 IQQR\ ? 5 , FGGG\ & + # s - 3 + s 1 8 , 5 KH V 1* + 5 U FG RG 5 * + 5 3 1 8 V , 5 V #$ V ) FGGK"# 8 V LWF 6+# 7 s - 3 1 , , 1, V , / 8 ) 8 5 1 1 FWF 6+ 5 & , 1 # , V 1 , / 8 / 8 8? 5 5 5 1 * + 58 s : 1 ) . ,, s 55 ,$ ! V ! 5 8 V 1 1 * + ! , 1 1 1 5 1 1 5 5 ) s 1 " V 1 1 , .8 FGGH"# . / . .8 1 , , ) .8 # !FGGH " 1 1 + / . FGGH " 8 1 1 #+ , 3 , 8 = IQRQ\ /8 ) !:2 55 ! 1 1 , s 1 1 !: 8 : : ,$:=FK" 5 IQQQ\ C ,8 FK 1 , 1 ? 5 ) / . , 8 1 1 !FGGH " 5 !C : 5 1 , # 1 j :, ) S 5 1 1 : 5 1 3 5 1 2 5 s s C IQQL" V1 1 1 1 , 18 ) 5 Meat Science accepted ! Capítulo VII IQQI\ # IQQH\ FGGG\ V 1 # FGGG" V1 1 55 V 1 8 ? .8 V / . , ) , 1 5 8 * + ( s IQQF\ s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s C / 8 1, V # IQQK\ IFQ 5 RG 5 * +# 1 ,$:=FK 1 , V ILG Jorge Ruiz Ramírez Tesis Doctoral 1 5 ) , # 1 V 1 ) 1 ) 1 V V . 1 , "# ;) V , U 5 1 1 8 , 5 5 1 ,1 F ' /$ 5"1 '#)8 !"'#)8 !(#%$ 8!#)8 !(#%$ B 8!#)8 !(#%$ 8!#)8 !(#%$ C S# 5 "# $FGS F S 1 5, ' 8! $V , % ] I I# V 1 U , 1 ! LL 5 G#GQ 1 o, FWF 6+ 5 JR WFa $ ) JG8JHa S 1 F WF 6+ ) # 8 V V V U F 1 FL 6+SRHa $ " # 1 V 1 - 5) V 1 1 5 !: +98 , FWF 6+ 5 ! 8 $FGS % F#M 1 V !: +98 o , "# , U , 1 FL 6+SRHa $" 1 5 1 , U , - , !KG×FG×FG f \ f ! '#7! +*#$#'5 /N1 /V 1 LL FWF 8 MI FWF JR#GWF#G IL IGWF MJ#HWF#H IIG IHWF HJ#HWF#H JF FRWF MH#GWH#G , V 1 ) V 1 , ! I"# $ V 1 V # !>$ 1 := ! C" V 1 , F V 5 !C 5 I"# 1 1 ×Q 1 , 1 , 8 5 , I# : 1 # , C% 5 :=% 8 # !&$ × !IR 1 f LFK C 1 , ,, ; 1 , ,, V " V 1 LWF 6+ HJ#HWF#Ha $ I S , Meat Science accepted 1 Capítulo VII ) ) V 5 HR#Ha V , 1 # V 1 FF#K 8 , U , , 1 G#Ia 9F ,1 ) 3 . 1 1 # :, V , " 8, U # 5 1 V V V ) , V 1 1 5 1 IQJR"# 1 1 1 ! V , ) 1 V , 5 I , V ! " , 1 ) 5"#" , V 5 8 IGLWF 6+ V 1 1 ! 9 + ! " V !0=" fG , 5 V 1 = ! *" 1 \ 5 1 ,1 8 # * + V 1 8 1 5 V S# ! 8 IQQG"# P !U $F9 S U 0="# 8 1 8 8 , 1 ! 8 ) , V 8 V 1 8 " 1 5 ?G >5"#&%&>!*#& # , HGa 5 1 % 1 t1 _! IG×IG×IG V V , 1 ) , "# C , V 8 , 1 5 1 1 1 , , , _ 1 8 5 1 1 5 1 5 ! U , 5 V 1 5 " V IQJR"# * , 1 8 1 SH :;= 5 ! ) , , 1 !7 ) ) 8 1 , 1 1 !D) 6 1 1 1 , # = : V 5 * + 1 _\ 1 t , 8 5V 5 1 , F±F 5 MG 1 5 G#I± U , 6+ 5 V , U 5 8 !: +98 o" V 1 QQ#R±G#Ia *F - t1 1 ) "# $ ILI 1 ! 8 IQJM"\ X( 1 8 :98IKKL !IQJL" 8, 8 ILF Jorge Ruiz Ramírez !* *" V 1 , Tesis Doctoral , 5, 1 !X IQRK" 5 1 = + ⋅ 5 1 1 V 1 1 #* + V , 1 M#FH" !I" 1, V V 5 , ! * + 1 , , !;3 8 I"# V 1 X( 5: : V 1 1 *- * 5 : : 5 , 8 !;3 F"# !0=8* + "# , ! " V , V # 1 , , IQQQ"# 1 V 1 C # 1 ) 1 55 # 1 0 , V ;3 . ;3 55 V : ,V , : :" 5 ) ) , # ! " V 1 5 V ) V 1 1 ; 8 >! , 8 ' , 8 2' ' ' 1 ' ,$c 5 1 5 C? !"+ '" )$ $#"&+""#%) 1, F# ! ; ,, 55 # V 1 , 8 V 1 5 1 ) 1 1 ,$:=FK * + V I ! -=" , , 1 C × !: : 1 5 1 # C , U = # # 1 V 1 1 : : 5 ) ) !? $ - , 1 1 V ) # 1 U $F9 S !U 0= # , ) 4 FGGF"# 1 V hU * +"V 1 1 s IQQL\ : 1 ) . ,, : !F" P1 * !+ . ! + 5 1 * * = ⋅ = ⋅ 8 1 Meat Science accepted ,$c! * + Capítulo VII ) " V 5 ,1 1 5 * , 3 1 1 V ,, 5 V 1 ) 5 ,1 , ,$:=FK * + # L 1 V ) 1 V 1 V 1 5 1 LLWKa# V * + V 1 V , - !+ s ? 5 .8 / . 3 * + V 5 1 5 1 C 1 !IQQH" ) = !IQQJ"# 5 1 V := U8 1 , V 1 , U 5 FGGF\ = = X 1 U # FGGH " 1 , ) 1 1 3 5 1 1 * + 1 * + 5 * + 1 s: ) 1 8 \ + 55 C + FGGF\ V # , ) = 1 1 = V * + V 1 = ) := V := ) 1 1 5 * + 1 1 + 1 1 V V " V1 1 55 * + , 55 8 , ! C 1 1 * + * + V1 55 8 # # $ V ) V L" * + 1 1 ) V1 ) 1 ! 1 1 1 1 1 ) 1 ) "× 1 1 1 ) * + V 8 1 # V1 1 # 8 1 * + , , 1 55 1 1 V 8 ) # 1 5 5 5 , , * + V * + V ! 1 ) # $ V ) 8 , ) 1 , 5 1 V ,$:=FK * + V 1 V C# 5 V # F := 1 55 5 V 1 1 ILL , 1 1 1 A. s C C FGGH"# = 1 := ! ) 1 ILK Jorge Ruiz Ramírez 1 1 1 1 IQQH"# 1 1 1 1 C 1 V1 ! 5 5 / 8 1 1 V C V 1 1 55 1 ) G#Q U 5 1 V 1 1 ,$:=FK 55 V C 1 , * * 1 1 5 V Z- ) 8 V 1 ,$:=FK ^ M#F# , ) 5 C C , 1 V 1 , 8 !IQRH" 1 = 4 C$# & ' ' 1 V 1 V # $ ) V V1 , 8 5 C , # !FGGH" 1 I#LG U $F9SU 0=# FGGL" # IQQH" 1 ) , 1 # , := 1 ) , C# 5 := ! 1 1 1, 5 1 ) 8 + 1 F 1 1 , 5 1 5 ) ! 6 C 1 5 = F V ) s+ >! # # ) 1 ) G#KH 1 !IQQK"# C V : 1 + 1 V V 5 * 1 1 $ V ) 1 8 := 5 V 1 # !FGGH" 5 # # !IQQR" V1 0=# # !IQQH"# : * + 5 $F9SU , 1 ) , 1 V 8 1 , V 1 1 V C : 5 1 V , ) ) # !FGGG"# * ) 8 , P 1 := 1 1 5 V1 1 # $ # , , ) Tesis Doctoral ) ) $F9SU 0=# G#JH V1 8 C 1 ) V G#JH U Meat Science accepted Capítulo VII ILH JG $ $ !U " ! " MG HG KG LG FG IG G G#FG G#KH G#JG G#QH I#FG I#KH !" G#MG G#JG G#HG G#MG :, + 1 ) @ !U $F9SU 0=" G#HG G#LG G#KG :, G#KG G#FG G#LG G#IG G#FG G#FG G#FG G#KH G#JG G#QH @ C F# $ 1 ! C" , % !=" := !\" C "! " , 1 I#FG I#KH G#KH G#JG ### ) 5 ) , !:=" ) , G#QH I#FG I#KH ! " ) !888" , ! V 8 1 1 " ! "# !@" 5 #; , ! N N ILM Jorge Ruiz Ramírez C L 1 V 1 1, , V ,$:=FK 1 1 5 V 55 5 , ;1 1 1 ) , , 5 V 1 " 8 ! 1 V 1 V 1 55 1 51 V 1 55 * + !IM#Ha ) IQ#Ha 0= 8 >!> + KH 2( ' ' 6 5 V 1 # 1 1 1 * + 5 1 := 1 5 VV 1 1 ,V 1 1 , * + IR#Qa ) FK#Ra 0="# 5 ) V # !IQQJ" 5 # !FGGH " 55 5 # !IQQR" V1 V 5 1 5 , 1 1 / . 55 !1 , := "# : 1 1 1 ,$:=FK^M#F ! , .8 5 1, , ) := V 1 ,$:=FK ] H#J 1 V 55 1 2 s 5 ) 1 1 1 ) 1 V "# 1 ) C ) IQQK\ :2 IQRR" "# , := 5 A + 8 IQQL\ V * + V ,$# 5 5 1 1 ,$ 1 1 1 5 1 ! / 8 5 !FGGG"# V 1 1 , := !IQQL" 1 . s V 11 1 , := ) V ,$ V V 1 1 s :2 1 V 1 1 1 1 V 1 ,$:=FK ^ M#F# 1 1 1 1 1 := ,$:=FK ] H#J 1 1 5 ! , 8 V 1 , 1 , ,# $ V 8 1 5 C# 1 1 # ) ,$:=FK , V 1 , , V 1 V ) Tesis Doctoral 5 5 := , 8 $ V % = ⋅ ⋅ + ⋅ 2( + ⋅ 2( u % 1 %+ A 8( %* %)!)'" $ B Z Z G? / = 1 B Z = 5 8 ? $$!$ !7! 1 ? 1 5 1 , +"& ! ? %@ /Q ?R1 / , 1 #8>/P G? 1 %@ / N1 !$#+* / N1 $$!$ !7! [ +"& ! #8>/KN1 I#KHWG#HI I#KKWG#KK I#KHWG#HM I#KHWG#KH I#KFWG#KL I#RQWG#IG I#GGWG#IM *: I#FKWG#KG I#FKWG#LM I#LLWG#KR I#FFWG#LK I#IJWG#LG I#HRWG#IK G#RQWG#IK vv RM#FHWL#RG RH#QGWL#IF RH#LJWL#RM RH#RKWF#JH RJ#GIWL#MJ RK#GIWL#FQ RR#GKWF#FR *: IF#LFWL#FJ II#JFWL#QR IF#LMWK#GJ IF#IQWL#GQ II#HGWL#RH IK#IHWL#RL Q#RQWI#HJ *: B Z / 1 IL#LMWH#LG IF#MHWK#JF R#GGWF#MK IK#HRWL#HI IM#KKWK#GI IM#IIWK#LM Q#QGWL#LK v B Z / J 1 FK#IHWK#FG IQ#HMWK#MQ FL#FRWM#GI FI#LJWK#LJ FG#QGWK#LQ FH#KKWL#KI IR#FMWL#KK *: V !V 1 # 1) " 5 55 ! ] G#GH"# Capítulo VII = V 1 55 1 0= f 0 # * + !0=" f + * + f !U $F9SU 0="# f !U $F9SU 0=8U * + "# *: f ^G#GH# vv f ]G#GI# v f ]G#GH# Meat Science accepted F !±:#0" 5 ,1 = ILJ ILR Jorge Ruiz Ramírez Tesis Doctoral L * + V $$!$ !7! * + = V 1 55 I % % U $F9SU ) B $ IG#KI±G#HH H#HR±I#GF I#JH±G#GQ G#RQ±G#FL B $ IJ#QG±G#HG II#FM±G#MK I#HK±G#GM G#QI±G#GQ B $ FG#GK±I#MM IF#RK±I#FK I#KM±G#GK G#RQ±G#GM 1 5 55 # 1 5 + ⋅ 2( + ⋅ 2( ⋅ :, % = + ⋅ + ⋅ 2( .8+ , V1 1 , # / 8 1 5 1 1 1 V # + 5 =C V V 1 , V1 V 1 F#R J#Fa# C K 1 V , * + S 8 , 5 55 3 1, 5 1 ) ) # 1 V 55 55 U * + 55 V1 1 55 1 , 5 1 1 V 5 1 1 5 5 , ) V 1 1 1 , !FGGG" 5 ) 5 1 V 1 $ V ) + ) B , ! ] G#GH"# # % = + ⋅ 1 ) /N1B +"& ! K + 1 ± :#0" ! " != , 1 # 1 1 5 1 , ) 8 Meat Science accepted Capítulo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orge Ruiz Ramírez ! =:;" 1 F ! Tesis Doctoral 55 5 1 8 ) , , " 5 1 5 1 V 1 5 1 1 1 V1 V 1 1 , , V 1 , 1 1 1 1 . *'A . + , " 1 V 1 !FGGH " 1 5 .8 V 1 # 5 ) * + 1 , 1 1 , 8 " 1 ,$:=FK 1 8 / . 1 1 LG#JHa 5 1 1 1 # 1 %)& +"#%)" V1 1 55 V FM#IIa 5 FR#KKa"# ) ) FH#JJa 1 ! 1 V FF#GQa 1 # , 55 ) ! ) 1 # 1 IQ#QMa"\ 1 5 8 IR#MMa FL#JGa"\ , ) V IL#HKa 5 5 ! ) , 1 , , ) 5 1 IH#HIa"\ 1 FI#LIa ! ) 1 1 V ) V1 1 , ) !# # 1 1 /8 5 , ) ! V 1 V V 1 IR#FMa ! ) 1 ) 1 , 5 ) 55 1 ) # 1 1 1 1 , , ' . s IQQR" 1 1 1 5 V , V ) 5 , 5 55 5 1 5 5 , 5 1 VV 1 5 , #$ V ) 1 H 1 V 5 1 V # 5 1 , 5 5 5 ) 3 ) C # 1 1 5 1 1 , ) 1 8 ,, 1 1 ) V 1 5 HGa# # !IQQH" 5 8 1 ) , 1 1 1 V1 1 , K# # , 1 1 , 5 1 # : , , 1 V 1 1 , 1 55 V 1 V ) 1 V Meat Science accepted Capítulo VII IKI JG MG ! " !U " HG IH#HIa IQ#QMa FL#JGa FR#KKa KG $ $ LG FG IG G G#FG G#KH G#JG G#QH I#FG I#KH !U $F9SU 0=" G#MG G#JG G#HG G#MG G#KG IQ#QMa ) FL#JGa G#LG G#KG IH#HIa IQ#QMa FL#JGa FR#KKa :, + 1 G#HG IH#HIa FR#KKa G#FG G#LG G#IG G#FG G#KH G#JG G#QH I#FG I#KH G#FG G#FG G#KH !U $F9SU 0=" C K# 1 % % f 1 1 , u N N2( N N2(! 1 N2(N 4 1 G#JG G#QH I#FG I#KH !U $F9SU 0=" ) ! " 5 ) % , 8 f 1 @ N ) V # ) N2( N 2(× ! " 5 , 8 IKF Jorge Ruiz Ramírez Tesis Doctoral K = 1 1 ) N N N N2( N2(N N2( 5 5 1 N N2(N 5 N2( , # FIJ#K 8F#MQML 8G#GFHH 8G#GLML G#RK H#RF %>!"#7!)!"" G#GMIRH G#LJMJI G#GGRIR 8G#GGJLH G#MQ G#MQ (#)8#)!"" G#FIHML G#FFFQF G#GGLK 8 G#MM G#MM 5 1 \ ($)!"" /. 1 4 4 ) 2( f , F 55 5 \ =:; # # !U $F9SU V V "# V V1 1 1 ) 1 8 1 1 1 5 # 1 5 V 5 1 ! 8 ) C9* + 5 !, 5 "# C .8 1 , O 5 5 . % FGGG FGGGLLG" 5 , , V 1 85 , , 5 5: 5 1 V 1 C / ._ 7 ) # !:!(!)&!" ,$:=FK 5 5 1 * + ) 5 9 + !IQQG"# , , ' # !IQQI"# V 1 1 # V 1 1 :, 1 = 1 5 U V -FGGL8GKMIF 1 1 * 1 , , 7 ) C :, # !IQQL"# ( 7 $ 4 H4 L8Q# # !IQQR"# 5 : 5 , ( # &X)%@ !$8!*!)'"? 1 E # IH 1 ; ,, QLI8QLH# ( 8 , 8 O 8 # / 8 # # , 8 Meat Science accepted Capítulo VII JG JG ) MG ) f IH#HIa HG !U " ! " " ! KG KG LG $ $ LG FG $ FG IG IG G G#FG G#KH G#JG G#QH I#FG G G#FG I#KH G#KH @ !U $F9SU 0=" G#JG G#QH I#FG I#KH @ 0=" !U $F9SU JG JG ) MG ) f FL#JGa f FR#KKa MG HG !U! " " HG ! " f IQ#QMa MG HG KG LG $$ $ $ IKL FG IG KG LG FG IG G G#FG G#KH G#JG G#QH I#FG I#KH G G#FG G#KH G#JG G#QH I#FG I#KH @ !U $F9SU 0=" !U $F9SU 0=" C H# ; , , 1 ) V ! " 1 5 55 , , 1 ) %5 IL#HKa IR#FMa ! ) IH#HIa"\ 5 IR#MMa FI#LIa ! ) IQ#QMa"\ 5 FF#GQa FH#JJa ! ) FL#JGa" 5 FM#IIa LG#JHa ! ) FR#KKa"# N2( N2(N 1 % N N 4 V1 1 ) 5 1 ,# ( 8 / ( # 7 % O 9) ! $ % &&' ( #$) !,,# JI8RM"# ; # # !FGGG"# , 8 3 5 5 ) # (( $ * / # ,,# FJ8KG# # # s+ , # !FGGL"# ;55 5 1 ) 1 8 5 1 , 1 , ,, 5 8 1 8 5 ## $ MH IFJH8IFRG# # # s -# !IQQK"# 1 55 5 5 . IKK Jorge Ruiz Ramírez ,$ 1 5 5 V1 8 Tesis Doctoral , 5 1 s 1 5 C # * , ' $ + MM FJQ8FRF# # -# + #s # !IQQH"# 1 1 1 55 . 5 :; 8 1 , #+ ' 4 D ML8MQ# # -# s # !IQQJ"# ;55 5 , 8 1 1 5 5 8 1 5 1# * ' $ + , JK IQL8IQR# # -# = ( ;# s # !IQQF"# ;55 5 ,$ 1 5 1 # >-' ( #$) FFQ8FLF# # -# s :2 +# !IQQR"# 1 55 5 1 ,$ * + 1 ,8 ) 1 8 5 8 1 #* ' $ + , ?? LRJ8LQF# # # A. # s C # !FGGH"# 8 :, 8 1 1 5 # 8 1 8 V 8 # # $ MQ FGI8FGR# *# +# !IQJM"# 5 1 ' , #? ) ' ( # 0 * V 4 U# =# +# !IQJR"# , 5 # + ) ' 4 > ? MF8MM JF# 1 :# = # + =# + + C C C # !IQQK"# 8 5 5 5 1 8 1 ## $ LJ KKQ8KHM# U8 U =# = # s ? # !FGGF"# U 3 , 1 8 5 8 1 5 0 #* , $ RG QRR8QQM# =# = # # . -# #s # !IQQL"# : , , 8 1, 1 5 8 8 , 8 1 #* + $ HR QMR8QJF# : !IQQR"# , : # ) * ) FK FGGL 5 C : P % 1 ,%SS5 # S # , # !FGGI"# ; : % 5 8 ) #." , / ( % 0# E A 4?D H8IG# # :# s * # !IQRH"# ;) , 2 # # #." , 57 ) 7 , FH KGG8KGJ# / 8 # & 8? 5 # , #s- 3 + =# !IQQQ"# : , 8 , 5 :, 1 8 1 5 5 ) #+ ' MJ KFL8KFJ# / 8 # & 8? 5 # , #s- 3 + =# !FGGG"# ) 5 1 , 0 Meat Science accepted $ - :, 1 8 1 5 5 ) # # $ HM I8M# / 8 # / 8 # & 8? 5 # , # s - 3 + =# !FGGK"# 8 1, V ,$ 5 8 8 1 3 # # $ MJ MFH8MLF# -# # # s # !IQQQ"# 1 5 5 ,$ 1 , , 5 8 1 ## $ HF FMJ8FJL# :98IKKL !IQJL"# 0 5 5 # ) % 8 9 . 5 : 8 . # X # !IQRK"# # ' # +1 1 %; $ V - # = U # # # :# + :# *# = X 1 C# X# ;# # s : ) # P# !FGGF"# + 8 1 ,, 1 3 55 1, U 1 1 , # * , $ RG QQJ8IGGK# = # = ) # # = # + =# 0# s C# !IQQJ"# +1 1 8 1 ! 1 " , 8 55 5 1 1 1 3 ## $ KJ FQ8KJ# # # s : 18 ) . ,, +# !IQQK"# 1, V 1 , ) , , 8 1 5 5 ) ## $ 4 >- IIJ8IFF# A =# + # =# s :2 +# !IQQK"# 0 ) , 5 1 , $ ) 8 U ) V 8 55 5 , 5 Capítulo VII :, IKH 1 8 1 #+ ' KQ IH8FI# / . =# =# *'A . C# + # # ' . =# ;# s =# # !IQQR"# ;) 5 , ) 5 8 5 8 1 #* , # RH QGH8QIF# .8+ # # + ) # B #s / +# !FGGG"# ,, 5 8 8 1 55 5 5 , # + . '( LL QI8QH# .8 / . # : @# #s # !FGGH "# 5 5 V V ) 8 8 8 # # $ 4 MQ4 D H D D! .8 / . # # : @# s # !FGGH " 1, V V * + ,$ , 8 ## $ , # :2 +# =# # = 5 # s + # !IQRQ"# ;55 5 ,1 ,1 1 ) 5 , , # # $ 4 FH FKI8 FKJ# :2 +# =# s / 8 # # !IQQL"# + , 8 5 , 1 , ! 0" ) 8 1 , 5 1 ) 1 V1 , # * ' $ + , 4 MF JI8JH# : : !IQQQ"# : : R#GI# + *+% : : # : @# .8 / . # # s # !FGGH"# , 8 IKM Jorge Ruiz Ramírez 5 ! Tesis Doctoral 5 8 1 5 5 V ) V # # $ MQ FKQ8FHK# : 1 ) . ,, +# 0 =# * + -# # .. -# s # !FGGF"# 5 V , , 1 # # $ 4 MG JJ8RL# # C =# C . :# =# s 2 C# !IQQQ"# 2 3 55 , , , U V 5 8 1 # # $ 4 HI FHH8FMG# 2 C# s ; 1 0# !IQRR"# ; 5 1 , 0 $ ) 8 1 # $ FL I8J# 2 C# ;# s C # !IQQL"# + 1 , 0 $ ) 1 , 5 8 1 # * ' $ + , MF IHJ8 IMI# 2 C# s C =# !FGGG"# 1 5 . , 8 5 , U 3 # + ' MQ LRJ8LQH# # # : 1 ) . ,, +# +# :# s =# !IQQH"# : 3 5 8 1 55 1 , ) , #* + $ 4 MG IIRL8IIRM# # . Discusión Global de los Resultados B? ! &#4) $! Capítulo VIII IKQ ) &'#7#$ $ $! 8+ 5 $! &%)'!)#$% $! 8+ &%) %" . #: = !IQQF" , (9*!'(%" $! '!<'+( 4 3 ) - . , F !+ ,/ ) 3 I 5 1 " 2 5 , / 2 #; 3 ! " S ) ! "# - 5 ) 3 ( , 8 ; IG"# , L " - C , 1 5 # ' K !+ ,/ ) 8 . 2 # 8 1 IF 3 . , - I ( 8 - GQ d GM 8 2 B ! , 8 / 3 ) 8 ( # , 5 2 ) , , 2 . , ) 3 , / . 5 ; , ) ] G JG , 8 IQRM" !IQJG" 3 5 , , ,B GM ) ( 1 1 3 5 G J\ # , , 8 "# 2 . #; ( IL , !+ ,/ , . , !+ ,/ 5 5 8 . # , # , 5 . #; " , , /8 ) , 2 ! 5 8 , ) , , , 8 , ) 8 IHG Jorge Ruiz Ramírez Tesis Doctoral IFG C( - IGG =' C := C := ( KH / FRQ / MG 0 . ! " RG KG FG G GF GK GM GR IG + C !U SU " IM ) , 2 1 IK IR ! f U $F9 S U =:" IF# IG# = , IF . # . ) %$! % &%) +(!F /" 1 "'#& ,# #$ $ /" 1 %>!"#7#$ $ L IHHI 8R GQKF H IQQL G RQ J FQ G GGGI FM HRI 8MK QHJ LQ JMQI G JR F IL G GGGI G LQKK I GGKK G GK G GGGI 8 G LL %$! % &%) +(!F /" 1 "'#& ,# #$ $ /" 1 %>!"#7#$ $ ) f, 2 F f 5 =:; f ) G GGMF 8G GFRL G GRQJ G QM K KI G GGGI 8G GRRR 8G FRIQ G GQLM G QG I KH G GGGI G LRGM G IIIM 8 G KM G GK G GGGI \ ! 4 # # 2 # U $FGSU "# Discusión Global de los Resultados Capítulo VIII IHI G RG G JG G MG + 1 ) G HG G KG G LG G FG C( =' C := G IG C := ( KH / FRQ / G GG GF GK GM GR !U C , ( IG IK IM IR $F9SU =:" IL# ) * F$ 9KuJ$F9 ! , IF 1 ) # ; # FGGL" . ) 5 ' , ( , . * + 3 P G JH !- ( # !+ ,/ , ) IQRK"# * + , 5 5 3 2 3 , , , 5 / / 8, # FGGK"# / 5 , . , 1 1 ) 8 ) , 5 ! ' "# ; 5 , 1 5 !+ ,/ . " B 3 . 1 , L 3 IHF Jorge Ruiz Ramírez Tesis Doctoral G QG G RG G JG G MG G HG ; G KG G LG G FG C( =' C := G IG C := ( KH / FRQ / G GG GF GK GM GR IG IF IK IM IR !U $F9SU =:" C ' IK# ( , - ) / IG 2 ) # C IL IK ) " ( FRQ !+ ,/ " 1 / / KH / 1 , , B , ' # 3 / B 3 ) 2 #- ' 8 ( 8 IF + ,/ , 5 !+ ,/ , " ) ( 1 - 5 1 ) , !+ ,/ , . 1 \ 1 . # 2 ) # ) , , # / Discusión Global de los Resultados 3 ' , - ) / 2 , Capítulo VIII IHL 5 ( 3 1 KH / / IF , 8 ' . 3 / ) , 8 - ) H S 5 1 , 5 ) B # IQQH" 3 , ' I S# ) , 3 , . 1 ) / , L 7 ( GR , "# ) . 3 ' ( , , 2 / IG 1 # 8 2 . 5 O + ,/ * + ) , . , 8 ,$:=FK A , 2 2 , , 8 , " !+ ,/ 5 8 ) GM, !+ ,/ " , , , 5 ) # !+ ,/ , 8 , ) 8 2 + , 8 , / #- 2 , != 8 , 2 ) # ! / ) . 1 8 , ) # FGGL" S 5 8 # , . , . ) 2 5 ) ( 8 , # + 5 #: 5 ) 6# + ,/8 ( N ! , / + ,/ 1 ( , , N ) FRQ / # ; f ,/ 5 , . # ' " ( !+ ,/ !+ ,/ "# IHK Jorge Ruiz Ramírez ? Tesis Doctoral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iscusión Global de los Resultados Capítulo VIII IHH =' JG MG MG HG HG KG KG LG LG FG FG IG IG KH / 0 . ! " FRQ / JG G G FG G KH G JG G QH I FG I KH G G FG G MG G HG G HG G KG G KG G LG G LG G FG G FG G JG G QH I FG I KH G KH G JG G QH I FG I KH G KH G JG G QH I FG I KH + 1 ) G MG G KH ; G IG G FG G QH I FG I KH G IG G FG G JG G MG G MG G HG G HG G KG G KG G LG G LG IH# ' G JG G JG G FG G FG C G KH G FG G KH G JG G QH I FG I KH G FG !U $F9SU =:" , 2 ( ' # IHM Jorge Ruiz Ramírez Tesis Doctoral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iscusión Global de los Resultados Capítulo VIII IHJ JG MG HG . ! " KG 0 LG FG ,$:=FK ] H J IG G G FG ,$:=FK ^ M F G KH G JG G QH I FG I KH ! " !U $F9SU =:" JG JG . ! " $ B MG MG HG HG KG KG LG LG 0 ,$:=FK ^ M F FG FG IG IG ,$:=FK ^ M F ! " ,$:=FK ] H J G G FG G KH G JG G QH I FG I KH ,$:=FK ] H J G G FG G KH !U $F9SU =:" C IM# ,$:=FK ! " ' !U ! "( G JG G QH I FG I KH !U $F9SU =:" $F9SU =:" # . , 1 ' IHR Jorge Ruiz Ramírez Tesis Doctoral G MG G JG ,$:=FK ] H J G HG G MG G KG ,$:=FK ] H J ,$:=FK ^ M F G LG G KG ; + 1 ) G HG G FG ,$:=FK ^ M F G LG G IG GF G KH GJ G QH IF I KH G FG G FG G KH G JG G QH I FG I KH I FG I KH ! " G MG G MG $ B G HG G HG ,$:=FK ] H J G KG ,$:=FK ] H J + 1 ) G KG ,$:=FK ^ M F G LG G FG ! " G FG G IG G FG G KH G JG G QH I FG I KH G IG G FG G MG ; ,$:=FK ] H J G HG G HG G KG G KG ,$:=FK ^ M F G JG ,$:=FK ] H J ,$:=FK ^ M F G LG G LG G FG G FG G KH G JG G QH I FG I KH G FG G FG G KH G JG @ IJ# 1 G QH B $ G MG C , G KH G JG G JG ! " ,$:=FK ^ M F G LG ' ,$:=FK ! " !U ' $F9SU =:" ! "( 1 ) # G QH I FG I KH Discusión Global de los Resultados Capítulo VIII IHQ KH KH ,$:= ^ M F . ! " KG LH LH LG LG FH FH FG QHa * + 0 FG IG FG FMa * + FG IM JQa * + IH ,$:= ] H J KG IH KIa * + J LIa * + IH IG R KJa * + H H G G LH G HH G JH G QH I IH I LH G G LH G HH G JH !U $F9SU =:" C IR# G QH I IH I LH !U $F9SU =:" !U $F9SU =:" * + , ,$:=FK . , 1 ' ' # - ,$:= , . * + Fa A * + A , , ) , ,$:=FK ] H J# ; 3 .2 * + A 5 Ha 5 "# ; , # !IQQJ\ IQQR" 3 ! # IQQQ\ FGGG" 3 , ) U 5 . Ra * + IQQM"# - ' ) 8 ( ) 1 ^ * + 3 1 , / !X 3 # IQRJ" ) ! !IH KIa * + , , " / G GH" !:1 5 5 3 5 !J LIa * + IM Ha " ,$:=FK ] H J * + A FK Ra !FG FMa * + , . 3 5 3 " IQ H , = ) # 3 , 8 ) ,$:= ^ M F IR Q " ' 1 5 5 8 IMG Jorge Ruiz Ramírez Tesis Doctoral G MG G MG ,$:= ^ M F G HH G HG G HG FG QHa * + G KH ) G LH + 1 G LG G LG R KJa * + G FH G FG G IH G IH G JH G QH I IH J LIa * + G FH G FG G HH IH KIa * + G KG G LH G IG G LH FG FMa * + G KH IM JQa * + G KG ,$:= ] H J G HH I LH G IG G LH G HH G JH !U $F9SU =:" G JG G MG G HG ; G KH G JG G MG G HG G KG G LH G LH G LG G LG R KJa * + G FG G IH G IH G JH G QH I IH J LIa * + G FH G FG G HH IH KIa * + G KH G KG G IG G LH FG FMa * + G HH IM JQa * + G FH G IG G LH I LH G HH G JH !U $F9SU =:" C , IQ# 1 ' 3 . ) , # ; / 3 .2 ) 5 , / ,$ , / ) 1 !4 ) 5 8 * +# ; , ) IQRM" 3 I LH 1 ) * + , * + A - I IH # ) ! ^ G GH" , G QH !U $F9SU =:" !U $F9SU =:" ,$:=FK ' I LH ,$:= ] H J G MH FG QHa * + G HH I IH !U $F9SU =:" ,$:= ^ M F G MH G QH ,$:=FK , 5 ( . 5 8 GM '8 Discusión Global de los Resultados * + A 5 .2 , 3 ) GM . ( 2 - , , 5 8 1 8 ) # * + 1 A 5 ,$:=FK . , 8 , / # / 5 , ! 3 1 . # 0 / 8 2 * + * + 3 8 , ( # ) / , . 8 5 , ) , 3 A 5 / 3 8 * + 5 ) 5 , , ) , !IM Ia" 3 # !IQQJ"# !Q Qa" ! IG"# 3 , # !IQQH" ' * + IQQH\ , 8 * + , ' !IQQJ"# # 5 5 , IG" !FGGG"# IQ"# ( 3 #; . 3 8 5 .2 , , A 1 * + ' ; 2 ' ) ,$:=FK ^ M F !C / , 8 , 2 2 3 3 ) * , ) ! ] G GH" ' 5 ' , ) A 1 1 ' ,$# * + ) * + 3 3 * + # IQQJ"# - ( 5 , 5 3 8 , 2 Capítulo VIII IMI , , = A? # , :!&'% $! &%)'!)#$% $! 8( " #)'( *+"&+ ( 5 (%'!% #"#" "%,(! 3 5 %" $! 2)$#&! $! (! &#4) !)'(! (9*!'(%" $! '!<'+( ! # IQQH \ = ; 5 * + IMF !±0#;" ( 5 , 5/ )'#$ $ $! N N KN I FWG L I FWG L I MWG I G QWG I I HWG H I KWG K I HWG M I HWG H I KWG K I QWG I I GWG F RM LWL R RH QWL I RH KWL Q RH RWF R RJ GWL J RK GWL L RR GWF L IF LWL K II JWK G IF KWK I IF FWL I II HWL Q IK FWL R Q QWI M IL KWH L IF JWK J R GWF M IK MWL H IM KWK G IM IWK K Q QWL L J 1 FK FWK F IQ MWK J FL LWM G FI KWK K FG QWK K FH KWL K IR LWL K /N15 /N1 f U PG I LWG H (%'!2) /N1 = R E"&+ % I FWG K 5 B Z / Q I $#$ I FWG K $ [ ' , A %* %)!)'!" 83 / 5 $F9SU 5 =: f f 8* + ! =: f * f ) " 5 * + f , ** f 5 , ) Jorge Ruiz Ramírez II# = ! ]G GH"# f , ## Tesis Doctoral Discusión Global de los Resultados MG , 2 IH HIa ( IQ QMa HG FR KKa KG 8 ( FL JGa 3 5 LG 0 IML A JG . ! " Capítulo VIII ) 8 ' 8 , . FG ( IG G G FG # ; G KH G JG G QH I FG , ) .8 I KH + # !FGGG" ) G MG . G HG ( #: 8 G KG 8 ) IH HIa IQ QMa G LG 8 3 8 , 2 8 + 1 FL JGa FR KKa G FG 5 G IG G FG G KH G JG G QH I FG 5 ) 2 I KH , / # 3 G JG 5 G MG FR , G HG , , IH HIa G KG J Fa# + ) / IQ QMa !/ ; FL JGa G LG G FG G FG , FR KKa , 3 5 % G KH G JG G QH I FG I KH 0 . % ⋅ + ⋅(2 + ⋅ (2 u = ⋅ C FG# $F9SU =:" ' ( !U + 1 , 2 / # ) % = + ⋅ , ; + ⋅ (2 + ⋅ (2 ⋅ % " 8 IMK Jorge Ruiz Ramírez = + ⋅ + ⋅ (2 - C Tesis Doctoral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w8 , # * , $ JJ FMRH8FMQF# U # $# X 1 =# !FGGG"# 18 8 ) 5 w8 , # * , $ JR FLLM8FLKL# # + , # # !FGGL"# * + , 8 55 55 8 ) 1 M 5, U1 ## $ ML FQ8 LK# # + , # # !FGGK"# = 55 ) 1 Discusión Global de los Resultados 5 55 8 1 , # # MJ FGL8FGQ# $ # !IQRM"# C , , 8 5 1 5 1 # + ,# K# # + #; # 1 # 7 ) 5 7 # ILH8IQQ# X 1 =# !IQQM"# 1 5 1 1 . , 5 # # $ KL :IQL8 :FGI# # :# 1 # # : =# # !IQRK"# 0 8 , , 1 5 8 # * , + ' LF ILRF8ILRH# = 1 # : =# = 8 1 4# $# X V :# V =# !IQQH"# * 8 8 ) 5 !* " 1 # * * $ + $ ) ' KF KLM8KKI# = # = ) # # = # + =# 0# C# !IQQJ"# +1 1 8 1 ! 1 " , 8 55 5 1 1 8 1 3 # # $ KJ FQ8KJ# 9_$ # # 0# # U 0# # ) P# # !IQQJ"# 1 5 , 1 55 # # $ KJ IRJ8FIG# A =# + # =# :2 +# !IQQK"# 0 ) , 5 1 , $ ) 8 U ) V Capítulo VIII 55 :, $ 5 1 8 1 KQ IH8FI# *# $ 1U , #+ IMH 5 ' # !IQQH"# # ; ? . 8; , A # *# !IQRM"# C 1 # + $ ; 1 ) 1 + , # * V 4 U# FKM8LGF# :2 +# =# / 8 # # !IQQL"# + , 5 , 1 , ! 0" ) 8 1 , 5 1 ) 1 V1 , # * ' $ + , MF JI8JH# :1 # P ?# # 8 ) # !IQRJ"# : , , 5 ) , !1 , 8 ,1 1 " 8 55 * + + +F # + # M IQQ8FGJ# : 0# 4 4# !IQQF"# 1 3 5 5 , # % 1 +1 5 C $# # : 1V . # P# $ # ; # = 0 UU * V 4 U# MMQ8MRQ# # C =# C . :# 2 C# !IQQQ"# 2 3 55 , , , U V 5 8 1 # # $ HI FHH8FMG# 2 C# s ; 1 0# !IQRR"# ; 5 1 , 0 $ ) 8 1 # # $ FL I8J# +# . -# !IQJG"# +1 ) , , , 8 5, U U , # * + $ + IMM Jorge Ruiz Ramírez KH FKR8FKQ# .8+ # # + ) # B # / +# !FGGG"# ,, 5 1 55 5 5 , # + . '( 4 LL QI8QH# # # : 1) 8 . ,, +# : +# # !IQQH "# ;55 , 8 , 8 . , ,8 " JG FI8LI# #( Tesis Doctoral # . ,, +# : !IQQH "# : 5 8 + IIRM# 4 -# # s 5 ) $ # : 1) 8 +# =# 3 8 1 55 1 , )8 , # * $ MG IIRL8 4# # !IQRM"# ;55 ,$ , # * HI JJK8JRG# + . Conclusiones Capítulo IX IMQ # I# 0 ( H# - , 5 I KI U ) SU 2 1 1 , 5 , 8 3 / , 8 , ) , , ,B 1 S 5 ) , . ) , 8 # : , # # F# ; , M# - ) . , ,$:=FK ^ M F 3 / . ( , , , ) ,B , ) , 1 . 3 , , ( 1 3 , , 5 5 # 5 ' ' ( SU , / , R# - G MG U * + ,$:=FK ] H J , ) ) . ) A # ( 2 , ( , B # 2 1 L# ; , , J# - , 8 5 8 ' 1 8 8 ) . ) G HG 1 # ; ( / ) , ' ) 5 8 3 , 3 # K# - * + 3 . , , , . ( , # 1 ) , " , 5 , !L IJG Jorge Ruiz Ramírez Q# / Tesis Doctoral ( ( , , 5 , , 8 , 2 # IG# ; ,$ / 5 . ) 8 1 , 5 , I KI U , 5 5 # ) G FQ 2 8 . , 1 # : / SU , 2 . , ) , ( , ) , # 3 ? +#F= *2(!F ? !(( ? %+ ? () +? # + # ( + , (!"!)'! &%*+)#& &#4) "! (!"!)'4 !) ! ] &! !,( $% !) 9&!(!"= " I / = L$! * (F% #+ ;8IJIFI = %)8(!"% C1 +)$# ;, A $! *4)^ II Congreso Mundial del Jamón Anexos - ( , , 2 # , ) ' ( : 1 # , 0 / , , ( , : , # IQRH\ IQQM" ,$ ! , , IQQK" , ' ( 8 # ! I8 M" 8 # IQQR" / FJ ! # , 2 , ; , ,/, LcI HcI H / , 3 # IQQJ\ IQQK\ = 5 # IQQR\ # IQQR" ! , , , ! / K Ka# 3 , = . B !C # IQQR"# * * 8 28 IH 6+# ; 1 ) 3 ! " S ) , ! " ( MG MRa ! # # !FGGG" .8+ ( , IQJJ"# - B 8 , ( , 1 ) , / , 5 , , ( IQ QR KQ RIa 5 8 K HF , IK MMa# ) *F ! , ,6!'#7%" 5 8 ) 5 , : +9- IH 6+ , ; , ) ( / 5 B ) 8 3 1 / # , ( IJH 1 IK . o" , / / 5 # IJM Jorge Ruiz-Ramírez Tesis Doctoral 5 )9#"#" $! '!<'+( ( ! " ) 3 - 5 ) , 8 5 , ; 2 % 4( f ,/, IcIcI Z ) . Z u , "! IQJR" . : H - ( ( % 4( f f ( f ) , \ f \ f ) 8B 7X" ) ( Z x( \ 8B \ = 8 5 , . 8 @ !: # 5 \ 5 , 5 ! ) ( x( f # S# , HGa ! , , 5 "# ; , 2 )9#"#" $! &'#7#$ $ $! 8+ / 1 5 5 ( 5 5 . , ) U p+ , 8 , ! 8 , 8 5 SU ", IGLWF # ; 3 , * FH 6+ , ; , C , F ) G RF# I 8 ) 2 : : !IQQQ"# ; , 8 3 # - . 1 ) # - ( 3 , ) G JL )9#"#" !"' $2"'#&% . . ( . , ) 5 ! 9 + IQQG"# : . 5 5 1 , . 8 , ; P : ) ) &%)'!)#$% $! 8+ / 1 * ) #; ) . ) : # . -= , 3 , II Congreso Mundial del Jamón , Anexos ? !"+ ' $%" $! )9#"#" $! &%7 (# )F %$! % &%) , +(!F /" 1 =B C B C "'#&#$ $ KCA = C %>!"#7#$ $ GC R CR G A AB +(!F /" 1 =CA "'#&#$ $ %>!"#7#$ $ BL vf dG GH\ vv f d G GI\ vvv f *4) / #8)#:#& )&# 1 [[[ RL L RG A RC C %$! % &%) RR G B K C RR C d G GGI\ *: f 5 , #+ [[[ [ [[[ ) , ) , . ) , , , . 3 2 5 . G LH# ; 2 , 8 5 .8+ # !FGGG" , / 5 G QG 1 G JG 3 ) , , I LG G MG\ , ( ( ) .2 , 8 3 , ( . 3 8 5 , , . !+ ( 3 3 8 ) , 8 ) , 2 . 5 # , #; , 8 3 , , F ) 3 # - . KQ J U S G MK , IJJ ) 5 . # # FGGG"# ; ) . , 8 / 3 , , 5 B ) ( ; , 1 ( , , #; ) . 8 8 5 IJR Jorge Ruiz-Ramírez Tesis Doctoral HG HG I F L KG K H +(!F /" 1 +(!F /" 1 KG M LG FG G G G#M G#J G#R G#Q G#L G#H G#J G#Q G#L G#H G#J G#Q "'#&#$ $ G#J G#M G#J G#M G#K G#K G#M G#J G#R G#Q %>!"#7#$ $ G#MH G#HH I#I I#L G#MH G#HH G#KH G#KH G#LH G#LH G#M C G#J G#R I# G#L G#Q ) " G#H G#J G#Q I#I ! " ) . ! I ### ( 1 ) I#L ! \U ' M"# , 2 # . 1 ) 2 ( I#L G#H G#H ! I#I G#R G#R "'#&#$ $ FG IG IG %>!"#7#$ $ LG " 58 ) , , 8 , 5 , 3 / SU B II Congreso Mundial del Jamón , 2 Anexos , , 5 , 8 # ; ( 1 . , = 5 , + ! / -FGGG8GHIJ8 K8GK"# C + * / !C9* + " ) , FGGGFGGGLLG , % 5 , 0 # .8 / .# . # !IQQJ"# # : # C IQR# -# s # # JK% IQL8 IJQ # -# s :2 +# !IQQR"# # : # C # JJ% LRJ8 LQF# 9 + !IQQG"# 5 955 +1 #IH 1 ; ,, QLI8QLH# =#+# !IQJR"# C 1# LF% MF8MM# + , # # s # !FGGG"# = : # HH% FQI8FQH# C # * # :# s = =#+# !IQRH"# )# 3 / # # # FH !I"% IIJ8IFK# , -# !IQJJ"# # # 5 1 * # # : # 1 # s +1 # RI !I"%RQ8QM# -# # # s # !IQQM"# #: #C # JG% HFM8HLG# = / -# + # 3 # =# s # !IQQR"# = : # KQ% IKH8IHL# # # s : 1 ) . ,, +# !IQQK"# = : # LR% IIJ8IFF .8+ # # + ) # B #s / +# !FGGG"# = : # LL% QI8QH# : # : : !IQQQ"# 7 y R#I # + *+ 7#:# # # ? +#F= *2(!F !(( ? () + ? %+ ? # + # ( + , #+ ;8IJIFI = ;, A X Congreso Anual en Ciencia y Tecnología de los Alimentos ? +#F= ; *2(!F !(( , 5 ? () + ? ) Anexos %+ ? 1 ) 2 . 5 ) . 1 # , 5 ! 1 #: : :# B *- * , 1 "# ; 1 2 ) ) , ) . . , -= )B . )B , . , , 5 1 , 1 ) #; . !L " 3 # : ) 5 . ) f! N F N "# - G RQ G QM ) , # ) ) # ( , 2 . ) ) / 2 #: 1 #; , 5 , 1 # ; , 1 ) 1 3 2 . 2 : : , ) B IGcIGcL ) 1 # # ; L 1 . 1 , . " 1 1 S S ! 5 IRL 1 , ) = /02 !/0 1 ! W # .8 ( + / . # , -# ! S IJIFI = # "# + + :, International Congress of Meat Science and Technology !"&(# '#7! !* #(#& 1 Anexos IRJ *%$! %: #)"'(+*!)' > ($)!"" &> )8!" #) $(5=&+(!$ *+"& !" " :+)&'#%) %: @ '!(&%)'!)' )$ 1, V 1 V 1 , V , 1 1 55 5* + ,$ 1 1 ,# 1 5 1 V 1 1, V V 1 5 8 , 1 * + ,$ 55 # 1 , V U IR 1 # $ 5 5 1 1 1 ,$]H#J 1 ,$^M#F $ !:=" FK81 , !,$:="# := B ! C" V , 5 1 1 V 1 G#H 5 X*9L G#L 5 * *9F , V 1 FG HG RG 5* + , U 5 # 1 V ) , U V 1 . , FWF 6+ 5 KH # 1 5 , 5 1 V 1 , , ,, # 1 5 1 V , U 3 ,1 1 # . V 1 V1 1 V 5 1 , U 5 1 , # 1 , ,, , 5 V , V V , !0=" ! f U $F9 S U 0="# : 1 V ,$ V 1 5 1 # 1 V V 1 1 : : , U 1 1 8 ! 9+ *- *"# 1 1 5 5 V G#R I#L 1 1 ] G#M 1 1 # 1 C , 1 V1 5 := , 1 1V # 1 , 5 1 V 1 V ,$:= , 1 1 1 5 1 V 1 1 1 ,$# 1 * + 55 V 55 , 1 ,$:=# 1 * + 55 V ) , 5 1 V 1 V ,$:= 1 1 5 1 V 1 1 1 1 5 V G#M I#L , 1 5 ,$:=# 1 * + 5 Fa Ha 1 1 5 Ra# 1 , V 5 ,, , 5 1 1, V 1 V ! "# 1 1 5 1 V 55 1 ,$:= 1 * + # 1 * + 1 1 , ,$:=]H#J# ) V 1 G#M 1 1 5 V 1 * + ,$# X , V %1 V ,$ * + 8 # International Congress of Meat Science and Technology , ) &X8(%+)$ Anexos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orge Ruiz-Ramírez :# :, "# := ! C" 5 Tesis Doctoral V G#H 8 5 X*9L G#L 5 n , 1 % F#G !R#IQa IM#MJa U V , ) $F G S V 1 o , S , !*fLFK" V ,, 5 1 # 8 1 FL FWF 6+ ,1 1 ,, # V V 1 c V , V V , 1 $ I S , 5 - ) V 8 5 1 8 1" # S#$ , 8 ) V , 8 V 5 I 5 8 1 8 # V 1 , ,, , V 5 V 1 , U 5 , 8 HGa 5 1 1 V , ) 1 c 1 1# C 5 1 , IG c IG c IG 5 ; LGG" ) 1 # 1 , ! LWF 6+ HJ#HWF#Ha 8 1 # V 1 1 5 G#GI != = : SH :;= 5 , 8 $FGS FS ; 1, !7 ) 8 V o, 6+SRHa$ " 1 5 U , 5 1 !: +9- , . 5 " V , U % ] I 1 8, U , V . 1 # 1 "# 5 , # # 1 1 , !K F F , V :, 5 8 ,1 % F#M KH : +9- V 1 , FWF 6+ 5 8 IH±F 6+ 5 FL 6+ S RHa $" . SFK1\ , JGa * + 1 ) , N LG , U , !: +9- 1 F S S JG + U , # , \V "V 1 ) 5 , F 5 FI#FQa" !0=" , U !FG n :#-# 1 ) , V 1 FG HG 5 * + , V 8 5 * *9F , RG , , 8 1 1 V 1 FR#H 8 HQ#Ja# B , V V 5 International Congress of Meat Science and Technology IGLWF 6+ V 1 1 1 ! 9 + IQQG"# P 5 1 , V G#R , !0=" ! ! 9 + IQQG" 1 1 ,$ ! "V 1 ) # #% G#R 8 / . " .8 V 1 8 + 8 FGGL"# 1 ! 1 .8 1 9 / . 1 8 1 # 1 V 55 F ! " ! "V 1 1 V 8 ! V #C F! " 1 V 1 ,$ := !,$:="# 1 % 1 /B1 1 , 1 , f V , 5 1 5 ) # 1 5 I 1 V 1 1 1, 5 , # ; , 8 V1 V G#M 1 5 55 1 V 1 ,$:= V ! ^G#GH"# * ) V1 1 1 1 I#L# , 55 5 1 ^ G#J , 8 V V 1 1 1 ,$ G#M V !"+ '" )$ $#"&+""#%) V 1 1 1 C 1 ,$:= , 1 := 1 1V 1 , , C 1 , 9+ V # 1 # 1 1 1 1 V , V C 8 , 55 1, 5 ) 5 1 !: : 1 # 5 1 1 V 1 1 , U G#M , 5 1 V V 1 1 $ P1 ]G#M 5 # 1 5 # , ! , *- *"# V1 1 55 ! :9 IRKI8I% IQQM" V 1 1 1 V 1 IQQQ"# 1 V 1 1 : : 5 I#L 1 $F9 S U 0="# : 5 , f U Anexos IQI 1 1 1 , , 5 5 IQF Jorge Ruiz-Ramírez 1 V 1 1 Tesis Doctoral V ,$:= , # 1 V 5 1 ,$ 1 , 5 ,$:= V1 1 1 5 Ha U V , ) ) 1 V 1 1 V 1 V ,$# V 1 1 # !IQQR" 1 1 ,$ 1 1 5 ) 1 5 V 1 1 ,$ 1 5 1 1 ) ,$ 1 1 5 1 L "# 1 , 5 5 1 1 1 V1 5 1 ) V G#M 1 5 1 !IQQR" 5 5 1 * + 55 , 55 55 5 3 !I" 1 * + V 5 1 V 1 1 5 1 ! =:;" ! 5 8 55 , 5 ) 1 " 1 ! " * + 1 V 1 ,$:=# F 5 ,$:= , 1 * + 1 # 1 8 V 5 1 , V 5 1 V1 8 5 , ) 1 V ,$# 1 #: 1 1 V 1 V * + 1 1 * + 1 1 1 5 5 1 1 1 V1 1 , # IQRJ" V1 1 V !:1 V1 1 V 1 1 , ) V G#M 1 ) V 1 , 1 * + * + 1 1 , 1 5 1 1 V 1 1 1 * + V ,$:=# L 5 1 1 V # !C 5 * + ) ) 5 5 , 55 1 * + V1 V I#J# 1 # ! !IQQQ" V1 5 I#L# 55 ,, , 5 V G#M 1 V 1 1 1 V 1 1 Ra G#M 55 V1 1 5 Fa 5 1 , , * + 1 1 V , I# 1 1 V V 5 , V 1 1 1 ,$:= V ,$:= 5 V 1 1 1 ,$:= , 5 Fa * + =:; 5 Ha 5 * + 1 V 1 # V International Congress of Meat Science and Technology %)& +"#%)" , V ,, , 5 5 1 1, V V 1 ! "# 1 1 V 55 1 ,$:= 1 * + # 1 , 5 1 1 * + 1 ,$:=]H#J# V 1 ) G#M 1 1 5 V * + 1 ,$# &X)%@ !$8!*!)'" 1 :, 1 V 1 = ! , ( "# U V 1 5 : 1 -FGGG8GHIJ8 K8GK 1 1 1 : * 5 C 1 C9* + 5 5 5 ) . 5 8 1 .8 5 5 / . 5 +9*0;:8-7? 5 1 !:!(!)&!" 9 + !IQQG"# IH 1 ; ,, QLI8QLH# # Anexos IQL # -# s :2 +# !IQQR"# * $ + , ?? LRJ8LQF# # !IQQI"# 7 ) C ; , A# C # !FGGI"# . " , / ( % 0# E A 4?D H8IG# -# # # s # !IQQQ"# # $ HF FMJ8FJL# ?# : :# s : U $# $ 4 MK !M" !IQQR"# + ' QMQ8QJF# :9 IRKI8I% IQQM# 0 5 1 8 I% 1 1 # ) % 9 . 5 : . # = -# :2 1 . # =# : ( 2 # s - , . *# !FGGL"# (( # * / # +2 8; , A # .8 / . # : @# # s # !FGGL"# (( # # * / +2 8; , A # .8 / . # : @# # s # !FGGK"# # $ # *+% : : # !IQQQ"# -!@ ! + : : # :1 # P ?# # s ) # !IQRJ"# + # M IQQ8FGJ# # # : 1 ) . ,, +# +# :# s =# !IQQH"# * + $ 4 K@ !M" IIRL8IIRM# IQK Jorge Ruiz-Ramírez I# ; Tesis Doctoral 5, 5 ! "# V 1 8 51 F 0 8 ,$ * + ,$:=] H#J 8F#GQRH 8I#QRJQ G#RF G#RF L#JG L#RQ ,$^M#F Fa K#QQRG K#KGJI 8F#FGFF 8F#IHFQ G#RK G#RL L#KI F#IM Ha M#IHKJ 8I#QIKI G#RM L#FQ Ra R#JGHI Fa K#HLKG Ha Ra K#LIGH 8I#HQLR 8F#LFKH 8F#KRKM 8I#QJJH G#JQ G#JL G#QI G#RJ K#FR K#GR F#LH L#HF * + ,$:= ^ M#F , # =:; H#HIIQ M#GQGL ,$]H#J ) ) 5 1 H#RJJQ \ F 55 5 \ =:; 45 40 " 30 ! 35 25 $ 20 15 10 5 0 0 .3 5 0 .5 5 P C 1 I# $ \ V " , 1 ! f 0 .7 5 0 .9 5 1 .1 5 !U $F9 S U 0=" 5 ) , # % "# 1 .3 5 , ! " International Congress of Meat Science and Technology 45 45 40 40 35 35 30 30 25 25 $ 20 20 C 10 10 := 5 0 0 .3 5 5 0 .5 5 0 .7 5 P F# 1 ! " 1 .1 5 1 .3 5 0 0.35 0.55 P 0.95 1.15 1.35 !U $F9 S U 0=" ! " V " 0.75 ! " ,$:= ! " # 45 45 40 40 35 35 30 30 25 FI#FQa* + 20 IM#MJa * + 15 IM#MJa * + R#IQa * + 25 FI#FQa * + 20 $ 0 .9 5 !U $F9 S U 0=" ! " 8 15 R#IQa * + 10 10 5 5 0 0.35 P C ,$:= ] H#J ,$:= ^ M#F 15 15 C Anexos IQH 0.55 0.75 0.95 1.15 1.35 0 0.35 !U $F9 S U 0=" ! " P " V L# 1 V 1 ,$:= ] H#J ! " ,$:= ^ M#F ! "# 0.55 0.75 0.95 1.15 1.35 !U $F9 S U 0=" ! " * + 8 M A ? +#F= *2(!F ? !(( #+ + = ; , A# ( ? # # (!"!)'! &%*+)#& &#4) > "#$% &! ' $ %)8(!"% * 5% $! +)$# 1 $! %+ ? () + #7 ( + , z # ;8IJIFI ( "+ (!"!)' &#4) !) ! ] *4)^ ! &+ "! &! !,( (9!) !(+! = " I /BK= $! III Congreso Mundial del Jamón 7 Anexos , F H 0 , B 5 ( ! Ra * +# , 5 FRQ IQQR" , IQQ / 1 ( , 3 , , ' , 1 # - 5 8 := ) Q 8 KcFcF #: ' 1 , , 5 B 5 8 , 3 / 5 ) ,$ ) 5 ,$ ! " FF Ja 1 # ) ) 5 *F 6+ , # FWI , / 5 / LG / 3 #: . 0 ! , ,$ HH IR , MF MQ Q Q !:=" GL * *9F , I S 1 ) FK 1 ! , SU . !HGa 8 ) ! 9 + " . ! " # U X*9L , 5 ) "# : !,$:=FK"# C GH IcIcI , ' 5 ,/, ,$ HJ . HI Ia A : . L±F 6+ 1 5 , 5 8 IQJM"# ; / ) ( ( 1 !** S* "uIGG# - ,$ # X( ! 5 5 ( B 8 1 , IQRK" , 5 #: 3 !** " !X !* " , # !FGGG" ) ( . . , LLa# 0 " IQQG"# @ ( FGG Jorge Ruiz-Ramírez ! N N Tesis Doctoral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ongreso Mundial del Jamón Anexos RG G MG ,$:=FK ^ H J JG G HG MG G KG ) . ! " HG LG + 1 KG 0 FGI ,$:=FK ^ M#F ,$:=FK ] M F G LG FG G FG IG G G FG ,$:=FK ] H#J G KH G JG @ !U C I FG SU I# ;5 # G KH ' 5 , / ]G GH\ v; , ,$:=FK , , 3 , , AS *: *: * + A # ,$:=FK , HJ , 8 , 5 # , ) . C# 3 3 , ) ( ,$:=FK ^ M F 1 , , , ; ] / 5 3 ' 8 ( I $#$ KN IF RWI F IR IWL L ( ' I KH ( ( 5 * + ) := )'#$ $ $! N N H MWI G II LWG M IR RWK Q IR IWF M I FG " 1 PG IG IWK M IH RWF G , G QH SU . Q R Q JWL G FG QWF R ) G JG @ !U * + "# 2 = 5 G IG G FG " A (# , !" [ /N1 /N1 I KH ,$:= I# + W ) ! G QH 8 , 5 , , . , 5 #- , ; , ) = , ) ! ( 1 5 + -FGGL8GKMIF"# , / 8 FGF Jorge Ruiz-Ramírez Tesis Doctoral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z # , ;8IJIFI ( "+ (!"!)' &#4) !) ! ] *4)^ ! &+ "! &! !,( (9!) !(+! = " I /BK= $! III Congreso Mundial del Jamón - ) , , / 2 5 Anexos , !** S* "uIGG# - , 5 ) ,$ ! ( # ) 5 5 HR Fa 1 5 # ) ) 5 *F 6+ , FWI , / . ( . LG Ma 1 / , . L±F 6+ 1 5 * + IQQF"# ; ! " 5 , 5 / LG / 3 # #: . , 5 ! 0 , ,/, 5 F ) MR ) IR , ,$ ! ( % % A 1 H J" S ) . ! " Ra"# !:=" " ! 9 + IQQG"# . ( ( ! N N N(2 N(2N 3 / " 1 5 FRQ ! , SU FK 1 , 8 ) ' !,$:=FK"! - # U !F H ; ,$ 5 5 ) MF1 HH 1 * + I !HGa "# : . " IcIcI , , FGH ( 1 ) 2 N N N(2 ; C I ! N(2N "# N(2N LLa# 0 ' ! C" ( ) KcFcF Q . #: ' , !** " !X !* " , IQRK"\ B ! 1 ) , X( IQJM"# ; / . , # - , 2 1 ) 2 ! =:;" FGM Jorge Ruiz Ramírez Tesis Doctoral I# = /W0#;" # # +(!F /" 1 KF FQJ 8I HQF %>!"#7#$ $ 8G FGH I LKQ 5 , 2 * + C ! 5 ( : G GIL 8G GKJ 8G JLH G GIK 8G GLL F \ ' G GFI \ G RM G HQ I JH G GL ) 2 =:; # G MG LG IR RFa FH KHa LG JFa 0 ; , FH G HG FG ) IH + 1 0 . ! " G KG IR RFa FH KHa LG JFa 0 ; , G FG H G G FG G KH @ !U C G JG G QH I FG SU . (# , !" [ I KH 1 /W0#;" 6% I MWG F IH MWH L FJ FWK F , / ]G GH\ v ; , G IG G FG G KH " I# # F# = = 5 G LG IG ) G JG @ !U SU ) ' * + '% I MWG I IM MWK M FL LWL L , ,$:= ( G QH ' )'#$ $ $! N N I RWG H I HWG I IG KWG M IJ QWL H FJ RWF R FK MWL F * + # A I FG I KH " ( C ( I $#$ KN I HWG I FG GWI J FL QWL K 5 ) III Congreso Mundial del Jamón Anexos 5 FGJ # I# - , . , 5 , - # !IQQH"# 7 / ' ( / ( , , , . 1 # : ) # ( , 5 , 5 , , 2 ) , 5 , ; / ! IQQR" 3 3 / , . 1 -FGGL8GKMIF"# C + , - / ( 2 . # , + ! . 5 = # . , # ) , ( 1 8 * / ) !C9* + " , % FGGGFGGGLLG , ) 5 1 ) 2 , 5 # 5 , 8 0 8 / .# ) 3 . 3 ( , .8 5 9 + # ; 5 5 , # , ) ,$:= ( ,$ 3 ( 3 !IQQG"# 87: # ; -# = ( # !IQQF"# LR 1 + =: ;# FFQ8 FLF# , *# +# !IQJM"# J 1 1 + # * V 4 U8 ( 7: # ,$ , # !FGGH" ' .8 / . X # !IQRK"# = 1 # +1 1 %; 5, $ V FGR Jorge Ruiz Ramírez - # / . ' !IQQR" .8 / = +# C =# * A . C# + # . =# =# # ,,# = # RH QIF# . # : @ # # !FGGH"# + # # # : 1 ) . ,, +# =# !IQQH"# # : MG IIRL8IIRM# Tesis Doctoral . 0 + 1 ) FK LF LL LR KI KL ) K H IJ FM FQ LG HJ JL RJ IKQ IHG { 2 HQ MF 2 5 3 / HJ HQ MF MJ JI 2 , 5 LR { LM LR LQ KK HJ HR MF MK MJ JI JL RJ IGL IFH L KF B K H FH LL LK HJ HR MG MM JL RJ IGL IFH IKQ IHG IHI IHF IHL IHK IMI IMQ { 2 HQ MF MJ MR JI + IH 0 0 0 0 KI . FL FK KI A , HJ ML MK . H FL FK FM FJ LG LF LL LK LR KI KK HR JI JL RJ IGL IFH IKQ IHG IHI IHL IHK IHH IHM IHJ IHQ IMG IMI IML IMK IMQ ;5 8 8 H II FK LF LL HJ HR HQ MK IGL IFH IHF IHK IHH IMI IMF IML KG KI KF , , 2 + , / IHM IHQ IMG + , FM IHM + 1 ) H FH LL LR LQ KF HR JF JL RJ IGL IFH IHG IHI IHF IHL IHK IHM IHR IMG IMI IML IMK IMQ { { { { { / , , ' ,$:=FK IHK IMI A IHM IHK IMI FIG Jorge Ruiz-Ramírez L K H FR FQ LG LI LK LH MG MI JL RJ IGL IFH IKQ IHI IHL IMQ ; H FH LL LK LM LR LQ HR JF JL RJ IGL IFH IHF IHL IHK IHM IHR IMG IMI IML IMK IMQ ; ) IQ FG FI HJ HR MH MM MJ MQ JG Tesis Doctoral ; C 5 LQ KL C 3 5 ( FL { , FL {{ B FL {{ FK { { ,$ FK {{, , / FH {{ FM {, FJ {{ , 1 FJ {{ FQ {{ ) FQ {{ A FQ {{ , , LL C LK KI > , FM MR JI IHL IMI IML IMQ 1 5 IKQ , / 8, / IHI {5 FL { { , {, 3 $ ) FJ FR FQ LG HR MG MI MH MM MQ JG { , / , IJ { , / IJ { IR II L , II IF 5 FL KI IF MR _ X( 1 B - = = MG HJ RJ IHG IHI IHF IHL IMQ { 5 LQ KI KL IQ FK LG LH HR HQ IMI IMF IML { 2 MG ML MR JI 5 5 , RJ L H IL FF FL FH LH H LQ KL HQ JL RJ IHG IMQ =' H MH JI IGL IHG IHI IHF IHL IHM IHQ IMI IMQ =B ,/ LM =B 5 KG =B 5 LM KL =B ) LM =B LM Índice Alfabético =B LJ =B =B 3 / FII , , 2 IKQ , IL IJ IR LF LH HJ MQ JG KG KG : * + K H IH FL FM LI LF LL HJ HQ MG ML MH MQ JI JL IGL IFH IHI IHL IHM IHQ IMG IMI IMF IMQ * , K H IF FM { 2 MR JI * K FM HJ IMF { 2 ML MR JI 9 B FL FF FL LF LL KI IMI ,$ K H IK FK FH FM LL LH LJ HJ MK MH MM MR JI IGL IFH IHL IHK IHM IHJ IHR IHQ IMG IMI IMF IMQ { 2 HQ ML MR , / L IK FH FR LH IK FG FF FJ FR LL LK LH KF MH MQ JG JI / IF IL FF FL FK FH FQ LF LL ML IKQ IHI IHM IMF IJ IR IQ FF FM FR LF LK LM MR JI IHF IHL IMI IML IMK B IHM H IL IH IJ FF FJ FR LI LF LH MF MH MM IMH { , IH { IM : MM $ H II IK LL ML MK MH MM JG JI IGL IFH IHG IHI IHF IHK IHH IHM IHJ IHR IMI IMF IML IMQ , L IL IH FM FJ FR FQ LG LM KG HQ MG MI MF MM JG { IH IM MQ { , IJ MQ { IR IQ JG L K H IK IQ FI FF LK LJ KG KF KL JI IKQ IHL { 5 FI { ( K H FI FF FL FK FH FM FR LG LI LF LL LK LH LQ KI HJ HR JG JL IFH IKQ IHG IHI IHF IHK IHH IHM IMI IML IMK { MF RJ IHG IHI IHI IHF IHK IHH IHM { ' MJ IGL IHG IKJ IKR IHG IHI { LM LJ LR KG HR MF MJ JG {, L FH FM LF LH LM { HQ MJ ) , 2 IKQ 7 LR LQ FIF Jorge Ruiz-Ramírez ML MM KG W P 8 . KK Tesis Doctoral