! " # !#$%"#" &'"$( " )* +, #$% &$- +.!/ 0#,( ! "# $! %!$ &! ! '! $ ( +1 %) *+(( 203 455 -- - 67 - 6 - 8 ) 77 - : - 78 - - L - 77 - -- - 9 7 - -8 )8 669 -8 66: -: 66: -; +6: -; .H .- 678 6: .. + 7 0)7 7 07 ;77 7- - 7 <7 - 78 7 78 <7 79 - 7 - 7L !%! 8. 6#! 8L ="#! L8 !#$%"#! -HH - 8 - -.8 - - -6- - -66 ))6: - -68 ; - -76 , - -76 , . -77 DEDICATORIA A mis queridsimos padres, Wilfredo y Yolanda, por su infinito amor y comprensin, a mi hermano Wilfredo y a todas aquellas personas por su apoyo incondicional, especialmente a Hamilton. A los cautivantes bosques nublados por su enigmtico encanto que insta a conocerlos ms y a respetarlos. /)0 12 (3 4 $5 ! " / - ! " # !#$%"#" &'"$( " )* +, #$% &$- +.!/ 0#,( . /)0 12 (3 4 $5 ! " / 67 67 Este trabajo es posible gracias a la oportunidad dada por el Mg. Asunción Cano y el Dr. Víctor Pacheco, quienes confiaron en mí para la realización de esta tesis. Debo agradecer en especial al Mg. Asunción Cano por acogerme en su laboratorio, orientarme y brindarme su apoyo durante el desarrollo de la tesis. Al Dr. Víctor Pacheco, investigador del Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, por hacerme partícipe del proyecto “Protección de la Biodiversidad en Bosques Montanos Fragmentados y Propuesta para Conservar el Bosque de Carpish, Huánuco”, ganador del Concurso Nacional por Proyecto de Investigación, “Santiago Antunez de Mayolo – 2001”, auspiciado por el Consejo Nacional de Ciencia y Tecnología (CONCYTEC) y mediante el cual se pudo financiar parcialmente la tesis. Agradezco de manera especial al Mg. Hamilton Beltrán del Herbario San Marcos (USM), por su gran e invalorable ayuda en el trabajo de campo y orientación durante el desarrollo de la tesis. Así mismo agradezco al Dr. John Clark y al Dr. Laurence Skog del Smithsonian Institution, Dr. James Smith del Boise State University y a la Dra. Blanca León del University of Texas in Austin, quienes me brindaron bibliografía especializada y ayudaron en la confirmación de las determinaciones, en particular al Dr. John Clark por ser mi maestro y afianzar mi interés en las gesneriáceas. Al Mg. Rodolfo Vásquez y a la Blga. Rocío Rojas del Jardín Botánico de Missouri por darme la oportunidad de poder iniciar mi investigación en botánica en los bosques nublados. A Marina Villalobos, Miguel Chocce y todos los compañeros de las expediciones por su amistad y hacer de las salidas de campo una experiencia gratificante. A Jorge Lingán, Marybel Morales y Maritza Rodríguez por ser estímulos constantes de superación y compartir su amistad. Agradezco a la Dra. Magda Chanco, a la Mg. María I. La Torre, a la Mg. Joaquina Albán, a mis compañeros de laboratorio, José Roque, Ángel Ramírez y Wilfredo Mendoza, y a Arturo Granda por sus valiosos comentarios y sugerencias. Doy un agradecimiento especial a los pobladores de la Comunidad de San Pedro de Carpish por su hospitalidad, confianza y prestancia para ayudar durante las expediciones científicas. Un infinito agradecimiento a mis padres por apoyarme a conseguir mis metas, es de ellos también esta tesis, gracias Wilfredo y Yolanda. /)0 12 (3 4 $5 ! " / 6 ! " # !#$%"#" &'"$( " )* +, #$% &$- +.!/ 0#,( 7 /)0 12 (3 4 $5 ! " / Se presenta el estudio taxonómico del Orden Scrophulariales en los bosques montanos húmedos de Carpish, departamento de Huánuco. Se realizó colectas entre los 1700 a 3100 m de altitud bajo técnicas estándares de estudios florísticos. El estudio taxonómico consistió en la elaboración de claves para familias, géneros y especies, y en la descripción, análisis taxonómico y distribución geográfica para cada especie. Se reportan 4 familias, 20 géneros y 38 especies; la familia más diversa es Gesneriaceae con 21 especies en 8 géneros, seguida de Scrophulariaceae con 12 especies en 8 géneros; Acanthaceae con 3 especies en 3 géneros y Lentibulariaceae con 2 especies en 1 género. Se han dado cambios nomenclaturales en algunas especies, con excepción en Lentibulariaceae. El género más diverso es Alloplectus, de la familia Gesneriaceae, con 8 especies. Las especies A. carpishensis, A. ichthyoderma y Utricularia unifolia, esta última de la familia Lentibulariaceae, tienen mayor rango altitudinal (hasta los 3000 m). A. carpishensis es una especie nueva para la ciencia. De las especies estudiadas en los bosques de Carpish, 9 son endémicas del Perú, la mayoría (5) pertenecen a la familia Gesneriaceae. Las especies A. dodsonii y Columnea fuscihirta, de la misma familia, son registros nuevos para la flora peruana. Diez especies amplían su rango de distribución al departamento de Huánuco. /)0 12 (3 4 $5 ! " / 8 ! " # !#$%"#" &'"$( " )* +, #$% &$- +.!/ 0#,( 9 /)0 12 (3 4 $5 ! " / ) 77 ) 77 A taxonomic study of the Scrophulariales is presented in the humid montane forests of Carpish, Huanuco department. The collects were realized between 1700 and 3100 m, following standard techniques of floristic studies. The taxonomic study consisted in the elaboration of keys for families, genera and species, and description, taxonomic analysis and geographic distribution for each one of the species. Four families, 20 genus and 38 species are reported; the most diverse family is Gesneriaceae with 21 species in 8 genus, followed by Scrophulariaceae with 12 species in 8 genus; Acanthaceae with 3 species in 3 genus and Lentibulariaceae with 2 species in 1 genus. Nomenclatural changes have been given in some species, with exception in the family Lentibulariaceae. The most diverse genus is Alloplectus, of the family Gesneriaceae, with 8 species. The species A. carpishensis, A. ichthyoderma and Utricularia unifolia, this last of the family Lentibulariaceae, can inhabit to more altitudes (until 3000 m). A. carpishensis is a new species to the science. Of the species studied in the forests of Carpish, 9 are endemic of the Peru, the majority (5) belong to the family Gesneriaceae. The species A. dodsonii and Columnea fuscihirta, of the same family, are new registrations for the Peruvian flora. Ten species enlarge their distribution range to the department of Huanuco. /)0 12 (3 4 $5 ! " / : ! " # !#$%"#" &'"$( " )* +, #$% &$- +.!/ 0#,( ; /)0 12 (3 4 $5 ! " / - 78 - 78 2< " 5 0! " $5 != " 2< 0!! 5>0" " >? < @ !"! ) " " ! " !" 1 < '! $! ""? ! '02 0!" ? $0 ! " A! ! $!B$ &! $0 " C 2)! 01 " B "" $ ! ' 0!! !? 0'!" < $$ " -;D " B$ ' !" ! EF!) G &!? .HH-I ) ! ))' $ 0!! 2 $ "! $ 0 2$? 5$! 2 2$ 0 ! " ! 2 01 $ $! !0 $ !"J0 $? "2 " " B "" " 52 < " ! ! ' B0! <K? "$ BA " $02 " !? $ 0' $ 1 "&) $ $C "$ ? 5)! 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ÍGUEZ, T.S. SCHULENBERG, V. PACHECO, P. de RHAM, H. ORTEGA, J.P. O’NEILL, D.F. LANE, H. BELTRÁN, M. HIDALGO, L. ARIAS, J. PEREZ, C. ALBÚJAR & T. VALQUI. 2001. 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