DIVERSIDAD DE LA FAMILIA POACEAE (GRAMINEAE) EN LA

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DIVERSITY OF THE FAMILY POACEAE (GRAMINEAE) IN THE REGION OF
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the collecting intensity and species richness were made. To assess the completeness of the inventory a rarefaction curve
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de especies y de la distribución de las especies
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del esfuerzo de recolección se consideraron
las muestras realizadas en el tiempo (especies
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estimada se calculó utilizando dos tipos de
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Doscientas treinta y cuatro especies y 11 taxa
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P
P
P
N
I
I
P
P
P
P
P
P
P
N
E
N
E
N
Subfam. Bambusoideae
Tribu Bambuseae
Chusquea
4
Ch. culeou E. Desv.
5
Ch. cumingii Nees
6
Ch. montana Phil.
7
Ch. quila Kunth
8
Ch. valdiviensis E.Desv.
P
P
P
N
I
N
Subfam. Arundinoideae
Tribu Arundineae
Phragmites
3
Ph. australis (Cav.) Trin. ex Steud.
Subfam. Chloridoideae
Tribu Cynodonteae
Cynodon
9
&DI¿QLV Caro & E.A. Sánchez
10 C. dactylon (L.) Pers.
Distichlis
11 D. scoparia (Nees ex Kunth) Arechav.
12 D. spicata (L.) Greene
Eleusine
13 E. tristachya (Lam.) Lam.
N
N
OR CV
Subfam. Aristidoideae
Tribu Aristideae
Aristida
1
A. pallens Cav.
2
A. spegazzinii Arechav.
TAXÓN
BCÑ
AC
ABCÑ
Ñ
CÑ
ABÑ
Ñ
Ñ
ABCÑ
AÑ
BCÑ
BCÑ
BÑ
PROV
ARG, BOL
ARG
ARG, BOL, PER
ARG
ARG, BOL, PER
ARG
0
ARG
ARG
ARG, BOL, PER
ARG
ARG
PAISES LIM
10
5
5
120
5
10
10
1290
5
10
15
25
64
105
30
280
120
200
2060
1600
2280
1600
800
265
570
1000
ALT
L inf L sup
5
7
21
1
4
55
12
2
33
14
6
28
8
NC
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
DISTRIBUCION EN CHLE
AP TA AN AT CO VA ME OH MA BB AR LR
x
x
x
x
x
x
x
x
LL AI MG
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I
Subfam. Ehrhartoideae
Tribu Oryzeae
Oryza
34 O. sativa L.
Subfam. Panicoideae
Tribu Andropogoneae
Bothriochloa
35 B. laguroides (DC) Herter
36 B. saccharoides (Sw.) Rydb.
Imperata
37 I. condensata Steud.
Schizachyrium
38 S. sanguineum (Retz.) Alston
39 S. spicatum (Spreng.) Herter
P
P
P
P
P
P
N
N
N
E
N
N
P
P
P
P
P
N
N
N
N
N
A
P
P
P
P
P
P
P
I
E
N
E
N
N
N
P
N
P
P
P
P
P
A
A
P
N
I
N
N
N
N
N
N
P
I
OR CV
Subfam. Danthonioideae
Tribu Danthonieae
Cortaderia
20 C. araucana Stapf.
21 C. pilosa (d’Urv.) Hack.
22 C. rudiuscula Stapf.
23 C. selloana (Schult. & Schult.) Asch. & Graebn.
24 C. speciosa (Nees & Meyen) Stapf.
Danthonia
25 D. araucana Phil.
26 D. californica Boland. var. americana (Scribn.) Hitchc.
27a D. chilensis E.Desv. var. aureofulva (E.Desv.) C.M. Baeza
27b D. chilensis E.Desv. var. chilensis
27c D. chilensis E.Desv. var. JODEULÀRUD Nicora
28 D. malacantha (Steud.) Pilger
Rytidosperma
29 R. lechleri Steud.
30a R. pictum (Nees & Meyen) Nicora var. pictum
30b R. pictum var. bimucronatum Nicora
31 R. quirihuense Baeza
32 R. violaceum (E.Desv.) Nicora
33 R. virescens (E.Desv.) Nicora var. virescens
Eragrostis
14 E. curvula (Schrad.) Nees
15 E. mexicana (Hornem.) Link subsp. virescens (J. Presl)
S.D. Koch & Sánchez Vega
16 E. pilosa (L.) P. Beauv.
17 E. polytricha Nees
Spartina
18 6GHQVLÀRUD Brongn.
Sporobolus
19 S. indicus (L.) R. Br.
TAXÓN
B
B
BCÑ
AÑ
BC
Ñ
ABÑ
BÑ
B
Ñ
BÑ
BÑ
ABCÑ
Ñ
ABCÑ
ABCÑ
C
ABCÑ
Ñ
ABÑ
Ñ
ABCÑ
B
ABCÑ
AC
BCÑ
Ñ
ABCÑ
BCÑ
PROV
ARG, BOL, PER
ARG, BOL
ARG, BOL
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG
ARG
ARG
ARG
ARG
ARG
ARG
ARG
ARG, PER
ARG
ARG, BOL
ARG, BOL, PER
ARG
ARG, BOL
ARG, BOL, PER
ARG, BOL
ARG
PAISES LIM
750
1400
30
10
10
125
800
1000
1550
355
820
1000
10
60
5
5
60
5
800
640
650
10
1145
10
4
20
10
10
100
750
1400
1090
180
740
175
1900
2200
1550
390
2000
2100
930
125
355
820
60
820
2000
2000
2000
2100
1450
1250
25
450
250
1000
120
ALT
L inf L sup
2
1
35
11
2
3
8
19
2
2
8
12
17
5
42
63
1
36
10
7
3
4
3
27
11
13
2
16
6
NC
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
DISTRIBUCION EN CHLE
AP TA AN AT CO VA ME OH MA BB AR LR
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
LL AI MG
3RDFHDHGHOD5HJLyQGHO%tR%tR&KLOHFINOT9/ET AL.
A
A
A
A
A
A
A
A
A
P
P
P
P
P
P
P
P
P
A
A
A
P
N
I
I
I
I
I
I
N
I
N
N
N
N
I
N
N
I
N
I
I
I
I
I
N
N
N
I
I
Subfam. Pooideae
Tribu Brachypodieae
Brachypodium
64 B. distachyon (L.) P.Beauv.
Tribu Bromeae
Bromus
65 B. berteroanus Colla
66a B. catharticus Vahl. var. catharticus
66b B. catharticus var. elata (E.Desv.) Planchuelo
67 B. erectus Huds.
68 B. hordeaceus L.
P
P
P
P
A
A
P
A
P
N
I
I
OR CV
Tribu Paniceae
Dichanthelium
42 D. sabulorum (Lam.) Gould & C.A. Clark
Digitaria
43 D. aequiglumis (Hack. et Arechav.) Parodi
44 D. sanguinalis (L.) Scop.
Echinochloa
45 E. colona (L.) Link
46a E. crusgalli (L.) P. Beauv. var. crusgalli
46b E. crusgalli var. mitis (Pursh) Peterm.
47 E. crus-pavonis (Kunth) Schult.
Panicum
48 P. capillare L.
49 3GLFKRWRPLÀRUXP Michx.
50 P. miliaceum L.
51 P. racemosum (P.Beauv.) Spreng.
52 P. urvilleanum Kunth
Paspalum
53 P. dasypleurum Kunze ex E.Desv.
54 P. dilatatum Poir.
55 P. distichum L.
56 P. urvillei Steud.
57 P. vaginatum Sw.
Pennisetum
58 P. glaucum (L.) R. Br.
Setaria
59 6SDUYLÀRUD (Poir.) Kerguélen
60 S. pumila (Poir.) Roem. et Schult.
61 S. verticillata (L.) P.Beauv.
62 S. viridis (L.) P.Beauv.
Stenotaphrum
63 S. secundatum (Walter) Kuntze
Sorghum
40 S. bicolor (L.) Moench
41 S. halepense (L.) Pers.
TAXÓN
BCÑ
BCÑ
ABCÑ
C
ABCÑ
B
C
ACÑ
BÑ
CÑ
CÑ
Ñ
ABCÑ
Ñ
ACÑ
Ñ
BCÑ
Ñ
Ñ
C
BCÑ
Ñ
CÑ
Ñ
CÑ
BÑ
CÑ
AC
Ñ
BÑ
PROV
ARG, BOL, PER
ARG, BOL, PER
ARG, PER
ARG
ARG
ARG, PER
ARG, BOL, PER
ARG, BOL, PER
ARG, BOL
ARG, BOL, PER
ARG, BOL
ARG, BOL
ARG
ARG, BOL, PER
ARG, BOL, PER
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG, BOL, PER
ARG
ARG
ARG, PER
ARG, BOL, PER
ARG, BOL
ARG
ARG, BOL, PER
ARG, BOL
ARG
ARG, BOL
ARG, BOL, PER
ARG, BOL, PER
PAISES LIM
5
10
5
10
5
125
5
20
120
10
10
240
10
200
5
25
-
10
60
165
5
5
120
10
10
60
10
30
120
60
390
2100
1820
10
800
125
30
300
280
120
280
240
950
200
2000
25
-
2000
60
165
20
140
250
300
950
2000
250
125
120
250
ALT
L inf L sup
24
14
13
2
67
1
6
8
4
2
5
2
29
1
15
1
1
21
1
1
14
37
2
15
1
15
10
8
8
1
5
NC
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
DISTRIBUCION EN CHLE
AP TA AN AT CO VA ME OH MA BB AR LR
x
x
x
x
x
x
x
x
LL AI MG
*D\DQD%RW
B. lithobius Trin.
B. madritensis L.
B. rigidus Roth
B. scoparius L.
B. secalinus L.
B. setifolius J.S. Presl
B. squarrosus L.
B. sterilis L.
B. tectorum L.
B. tunicatus Phil.
Tribu Poeae
Agrostis
82 A. arvensis Phil.
83 A. brachyathera Steud.
84 A. breviculmis Hitchc.
85 A. capillaris L
86 A. castellana Boiss. et Reut.
87 A. gigantea Roth
88 A. glabra (J.Presl.) Kunth var. melanthes (Phil.)
Rúgolo & De Paula
89 A. idahoensis Nash
90 A. imberbis Phil.
91 A. inconspicua Kunze ex E. Desv.
92 A. leptotricha E.Desv.
93 A. mertensii Trin.
94 A. meyenii Trin.
95 A. perennans (Walter) Tuck.
96 A. philippiana Rúgolo & De Paula
97 A. scabra Willd.
98a A. stolonifera L. var. palustris (Huds.) Farw.
98b A. stolonifera L. var. stolonifera
99 A. uliginosa Phil.
100 A. umbellata Colla
Aira
101 A. caryophyllea L.
Alopecurus
102 A. magellanicus Lam.
Amphibromus
103 A. scabrivalvis (Trin.) Swallen
Anthoxanthum
104 A. altissimum (Steud.) Veldkamp
105 A. gunckelii (Parodi) Veldkamp
Tribu Meliceae
Glyceria
79 *ÀXLWDQV (L.) R.Br.
Melica
80 M. commersonii Nees ex Steud.
81 M. violacea Cav.
69
70
71
72
73
74
75
76
77
78
TAXÓN
P
P
P
P
P
P
P
P
P
P
P
P
P
P
P
P
P
P
P
A/P
A
P
P
P
P
N
N
N
N
N
N
N
N
N
N
I
I
N
E
I
N
N
E
N
P
P
E
E
E
N
N
I
I
I
P
P
A
A
A
A
P
A
A
A
P
I
N
I
I
I
I
N
I
I
I
N
OR CV
C
AÑ
BCÑ
ACÑ
ABCÑ
B
AÑ
ABÑ
ABCÑ
ABCÑ
BÑ
BÑ
BÑ
BCÑ
AC
ABÑ
ABCÑ
Ñ
C
C
BÑ
B
ABCÑ
ACÑ
AÑ
ACÑ
ABCÑ
C
ABCÑ
B
ABCÑ
CÑ
A
BÑ
B
CÑ
B
ABÑ
PROV
ARG
ARG, BOL, PER
BOL, PER
ARG
ARG
ARG
ARG, BOL, PER
ARG
ARG
ARG, BOL, PER
ARG, BOL
ARG, BOL, PER
ARG
ARG
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG
ARG, BOL, PER
ARG
ARG, BOL
ARG, BOL, PER
PER
ARG
ARG
ARG
ARG
ARG
ARG
ARG
ARG
PAISES LIM
5
90
10
5
5
2070
10
175
30
10
220
950
1000
100
10
70
5
450
260
10
2070
1735
5
10
10
20
5
10
5
1039
5
10
30
1000
390
10
790
5
5
280
300
125
2000
2070
120
2200
2000
2070
2200
2200
1600
2070
200
2070
2000
450
260
10
2200
1735
2000
1026
270
275
800
10
1250
1039
1030
250
780
2000
390
50
790
2500
ALT
L inf L sup
2
3
13
5
59
3
3
17
29
28
5
7
4
6
7
6
16
1
1
2
2
1
17
15
6
5
61
2
29
1
33
2
5
14
1
3
1
11
NC
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
DISTRIBUCION EN CHLE
AP TA AN AT CO VA ME OH MA BB AR LR
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
LL AI MG
3RDFHDHGHOD5HJLyQGHO%tR%tR&KLOHFINOT9/ET AL.
106 A. odoratum L.
107 A. redolens (Vahl) P. Royen
108 A. utriculatum (Ruiz & Pav.) Y. Schouten & Veldkamp
Arrhenatherum
109 A. elatius (L.) P. Beauv. ex J. et K. Presl ssp. bulbosus
(Willd.) Schüb. et Martens
Avena
110 A. barbata Pott ex Link
111 A. fatua L.
112 A. sativa L.
113 A. sterilis L.
114 A. strigosa Schreb.
Briza
115 B. maxima L.
116 B. minor L.
Bromidium
117 B. anomalum (Trin.) Doell ex Mart.
118 B. trisetoides (Steudel) Rúgolo
Calamagrostis
119a C. gayana (Steud.) Soreng var. gayana
119b C. gayana (Steud.) Soreng var. neuquenensis
(Rúgolo) Soreng
120a &YLULGLÀDYHVFHQV (Poir.) Steud. var. montevidensis
(Nees) Kämpf
120b &YLULGLÀDYHVFHQV (Poir.) Steud. var. YLULGLÀDYHVFHQV
121 C. viridis (Phil.) Soreng
Calotheca
122 C. brizoides (Lam.) Desv.
Catapodium
123 C. rigidum (L.) Dony
Chascolytrum
124 Ch. subaristatum (Lam.) Desv.
Corynephorus
125 C. divaricatus (Pourr.) Breistr.
Cynosurus
126 C. cristatus L.
127 C. echinatus L.
Dactylis
128 D. glomerata L.
Deschampsia
129 D. airiformis (Steud.) Bentham & Hook.
130 D. berteroana (Kunth) Trin.
131 D. cespitosa (L.) P.Beauv.
132 D. looseriana Parodi
Festuca
133 F. acantophylla E. Desv.
134 F. magellanica Lam.
135 F. monticola Phil.
136 F. purpurascens Banks & Soland. ex J.D. Hook.
TAXÓN
P
P
P
P
A
A
A
A
A
A
A
A
A
P
P
P
P
P
P
A
P
A
P
A
P
P
A
A
A
P
P
P
P
I
N
N
I
I
I
I
I
I
I
I
N
E
N
N
N
N
N
N
I
N
I
I
I
I
N
N
N
E
N
N
N
N
OR CV
Ñ
BÑ
BÑ
ACÑ
Ñ
CÑ
BCÑ
C
ABCÑ
CÑ
ABCÑ
C
ABCÑ
C
CÑ
C
C
C
Ñ
BÑ
BCÑ
CÑ
ABCÑ
ABCÑ
ABCÑ
CÑ
CÑ
Ñ
AÑ
ACÑ
AC
AÑ
ABCÑ
PROV
ARG
ARG
ARG
ARG
ARG
ARG
ARG, BOL
ARG, BOL, PER
ARG
ARG
ARG
ARG, BOL, PER
ARG, BOL
ARG
ARG, BOL, PER
ARG
ARG
ARG
ARG
ARG, PER
ARG
ARG, BOL, PER
ARG, BOL, PER
ARG, BOL, PER
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG
ARG, BOL
ARG, BOL
ARG
PAISES LIM
2000
1300
1720
10
60
10
30
10
10
10
10
10
5
10
10
100
10
10
2200
1800
10
150
5
5
10
10
10
70
10
5
20
10
2600
2000
1380
280
60
100
2000
10
1200
800
2000
10
1400
150
300
100
150
200
2200
2200
580
300
590
470
900
200
650
260
1030
180
280
650
ALT
L inf L sup
1
3
3
4
1
7
6
2
23
5
59
1
107
2
18
1
8
6
1
8
11
2
59
59
21
7
4
1
2
11
6
3
31
NC
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
DISTRIBUCION EN CHLE
AP TA AN AT CO VA ME OH MA BB AR LR
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
LL AI MG
*D\DQD%RW
137 F. scabriuscula Phil.
138 F. thermarum Phil.
139 F. tunicata E.Desv.
Gastridium
140 G. phleoides (Nees & Meyen) C.E. Hubb.
Gymnachne
141 G. koelerioides (Trin.) Parodi
Hainardia
142 H. cylindrica (Willd.) Greuter
Helictotrichon
143 H. bulbosum (Hitchc.) Parodi
Holcus
144 H. lanatus L.
Koeleria
145 K. fueguina C.E. Calderón ex Nicora
146 K. kurtzii Hack.
Lagurus
147 L. ovatus L.
Leptophyllochloa
148 L. micrathera (E.Desv.) C.E. Calderón
Lolium
149 /PXOWLÀRUXPLam.
150 L.perenne L.
151 L. rigidum Gaudin
152 L. temulentum L.
Nicoraepoa
153 N. andina (Trin.) Soreng & L.J. Gillespie
154 N. chonotica (Phil.) Soreng & L.J. Gillespie
Parapholis
155 P. incurva (L.) C.E. Hubb.
156 P. strigosa (Dumort.) C.E. Hubb.
Phalaris
157 P. amethystina Trin.
158 P. angusta Nees ex Trin.
159 P. aquatica L.
160 P. canariensis L.
Phleum
161 P. alpinum L
162 P. pratense L.
Poa
163 P. annua L.
164 P. bonariensis (Lam.) Kunth
165 P. cumingii Trin.
166 P. denudata Steud.
167 3S¿VWHUL Soreng
168 P. gayana E.Desv.
169 P. hachadoensis Nicora
170 P. holciformisJ.Presl.
171 P. lanuginosa Poir.
TAXÓN
P
P
P
A
P
A
P
A
P
P
A
P
A
P
A
A
P
P
A
A
A
A
P
A
P
P
A
P
P
P
P
P
P
P
P
N
N
E
I
E
I
N
I
N
N
I
N
I
I
I
I
N
N
I
I
E
N
I
I
N
I
I
N
E
N
E
N
N
N
N
OR CV
ABCÑ
B
C
ACÑ
B
Ñ
B
Ñ
BCÑ
CÑ
CÑ
ACÑ
CÑ
Ñ
AC
C
BÑ
Ñ
AC
ABCÑ
BCÑ
C
ACÑ
Ñ
B
ABCÑ
CÑ
CÑ
ABCÑ
ABCÑ
BÑ
ABÑ
Ñ
PROV
ARG
ARG
ARG
ARG
ARG
ARG, BOL, PER
ARG
ARG
ARG, PER
ARG, BOL, PER
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG
ARG
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG, BOL, PER
ARG
ARG
ARG
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG
ARG, PER
ARG
ARG
PAISES LIM
10
1200
5
10
200
1945
1430
1945
10
10
100
40
10
10
10
5
50
935
2200
20
5
5
10
10
5
1600
1700
10
10
80
10
10
1140
650
1500
1800
1200
20
1720
300
1945
1430
2000
1300
2300
650
250
125
350
250
5
50
1550
2200
285
800
280
650
30
150
1600
1700
2000
40
300
300
1500
1700
2060
1500
ALT
L inf L sup
19
8
16
18
1
5
2
2
15
13
4
5
2
11
6
2
1
5
1
32
16
9
4
2
32
1
1
72
4
3
22
38
6
21
1
NC
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
?
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
DISTRIBUCION EN CHLE
AP TA AN AT CO VA ME OH MA BB AR LR
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
LL AI MG
3RDFHDHGHOD5HJLyQGHO%tR%tR&KLOHFINOT9/ET AL.
Tribu Stipeae
Amelichloa
200 A. caudata (Trin.) Arriaga & Barkworth
201 A. brachychaeta (Godr.) Arriaga & Barkworth
Jarava
202 J. plumosa (Spreng.) S.W.L. Jacobs et J. Everett
Nassella
203 N. chilensis (Trin.) E.Desv.
204 N. duriuscula (Phil.) Barkworth
205 1¿OLFXOPLV(Delile) Barkworth
206 N. gibba (Phil.) Muñoz-Schick
172 P. obvallata Steud.
173 P. pratensis L.
174 P. resinulosa Nees ex Steud.
175 P. stenantha Trin.
176 P. trivialis L.
Podagrostis
177 3VHVTXLÀRUD(E. Desv.) Parodi ex Nicora
Polypogon
178 P. australis Brongn. ex Duperey
179 P. chilensis (Kunth) Pilg.
180 P. elongatus Kunth
181 P. exasperatus (Trin.) Renvoize
182 P. imberbis (Phil.) Johow
183 P. interruptus Kunth
184 P. linearis Trin.
185 P. maritimus Willd.
186 P. monspeliensis (L.) Desf.
187 P. viridis (Gouan) Breistr.
Relchela
188 R. panicoides Steud.
Rostraria
189 R. cristata (L.) Tzvelev
Schoenodorus
190 S. arundinaceus (Schreb.) Dumort.
Trisetum
191a T. barbinode Trin. var. barbinode
191b T. barbinode Trin. var. KLUWLÀRUXP (Hack.) Louis-Marie
192 T. caudulatum Trin.
193 T. nancaguense Finot
194 T. preslei (Kunth) E.Desv.
195 T. spicatum (L.) K. Richt.
Vulpia
196 V. antucensis Trin.
197 V. bromoides (L.) Gray
198 V. muralis (Kunth) Nees
199a V. muyros (L.) C.C. Gmel. var. megalura (Nutt.) Auquier
199b V. myuros (L.) C.C. Gmel. var. myuros
TAXÓN
P
P
P
P
P
P
A
A
A
A
A
N
N
N
E
N
N
N
I
I
I
I
P
P
P
P
P
I
N
E
N
E
A
I
P
P
N
N
P
A
P
P
P
P
A
A
A
P
N
N
N
N
N
N
E
I
I
I
P
P
P
N
N
N
P
P
P
P
P
N
I
N
N
I
OR CV
C
CÑ
ABCÑ
Ñ
BCÑ
BC
B
BCÑ
ACÑ
B
BCÑ
C
BÑ
Ñ
ABCÑ
C
Ñ
BCÑ
ABCÑ
Ñ
ABCÑ
ABCÑ
CÑ
Ñ
AC
C
Ñ
CÑ
Ñ
BCÑ
C
BÑ
BÑ
BCÑ
BÑ
Ñ
ACÑ
PROV
ARG
ARG
ARG
ARG
ARG, PER
ARG
ARG, PER
ARG, PER
ARG, BOL, PER
ARG
ARG
ARG, BOL, PER
ARG
ARG
ARG
ARG, BOL
ARG, PER
ARG
ARG
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG, BOL, PER
ARG
ARG
ARG, BOL, PER
ARG
ARG
ARG, BOL, PER
PAISES LIM
200
15
5
300
5
20
120
50
5
75
10
10
1000
2000
10
250
2000
140
10
10
10
5
20
20
10
10
100
20
64
10
5
1000
1300
5
930
2200
10
200
450
1100
300
800
100
120
700
670
930
650
10
2000
2000
1350
250
2000
2000
450
10
1350
2100
150
20
30
10
100
60
136
1200
150
2200
2400
2000
2100
2200
1430
ALT
L inf L sup
1
13
23
2
17
7
1
7
25
3
22
4
30
2
46
1
4
14
6
1
16
55
4
1
3
1
11
2
2
16
12
13
17
19
2
3
15
NC
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
DISTRIBUCION EN CHLE
AP TA AN AT CO VA ME OH MA BB AR LR
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
LL AI MG
*D\DQD%RW
Tribu Triticeae
Elymus
225 E. angulatus J. Presl
226 E. patagonicus Speg.
Hordeum
227 H. chilense Roem. & Schult.
228 H. comosum J. Presl
229 H. jubatum L.
230a H. marinum Huds.
230b H. marinum Huds. subsp. gussoneanum (Parl.) Thell.
231 H. murinum L.
232a +SXELÀRUXPHook.f.
232b +SXELÀRUXPsubsp. halophilum (Griseb.)
Baden & Bothmer
233 H. vulgare L.
Triticum
234 T. durum Desf.
x= Distribución conocida; ? = Presencia probable.
207 N. gigantea (Steud.) Muñoz-Schick
208 N. juncea Phil.
209 N. laevissima (Phil.) Brakworth
210 N. longiglumis (Phil.) Barkworth
211 N. macrathera (Phil.) Barkworth
212 N. manicata (E.Desv.) Barkworth
213 N. neesiana (Trin. et Rupr.) Barkworth
214 N. parodii (Matthei) Barkworth
215 1S¿VWHUL(Matthei) Barkworth
216 N. philippii (Steud.) Barkworth
217 N. poeppigiana (Trin. & Rupr.) Barkworth
Pappostipa
218 P. speciosa (Trin. & Rupr.) Romaschenko
Piptochaetium
219 P. angolense Phil.
220 P. bicolor (Vahl) E.Desv.
221 P. depresum (Steud.) C. Peña
222 P. panicoides (Lam.) E.Desv.
223 P. setosum (Trin.) Arechav.
224 P.stipoides (Trin. et Rupr.) Hack. et Arechav. var. stipoides
TAXÓN
P
P
P
A
A
A
P
P
A
A
N
N
N
I
I
I
N
N
I
I
P
P
P
P
P
P
E
N
E
N
N
N
P
P
P
N
N
N
P
P
P
P
P
P
P
P
P
P
P
N
E
N
N
E
N
N
E
E
N
N
OR CV
Ñ
B
C
ABCÑ
BCÑ
Ñ
ABCÑ
Ñ
ABCÑ
B
BCÑ
Ñ
BC
ABCÑ
ABCÑ
ABCÑ
CÑ
ABCÑ
BCÑ
ABCÑ
A
ABCÑ
Ñ
Ñ
BCÑ
ABCÑ
ACÑ
Ñ
Ñ
ABCÑ
PROV
ARG, BOL
ARG, BOL, PER
ARG, BOL, PER
ARG
ARG
ARG
ARG
ARG
ARG, BOL, PER
ARG
ARG
ARG, BOL, PER
ARG
ARG, BOL
ARG
ARG
ARG, BOL
ARG
ARG
ARG
PAISES LIM
300
1470
10
5
5
125
10
300
5
1380
10
1500
10
10
5
4
10
30
5
5
925
5
60
120
10
5
10
60
100
10
300
1470
10
120
2100
125
500
300
950
1380
2000
1500
120
650
1000
1400
650
300
1500
650
925
930
125
2000
300
1030
300
300
100
1400
ALT
L inf L sup
1
3069
1
2
19
17
1
9
4
19
1
9
1
2
10
34
29
9
32
7
61
1
50
2
2
22
45
9
13
2
24
NC
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
DISTRIBUCION EN CHLE
AP TA AN AT CO VA ME OH MA BB AR LR
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
LL AI MG
3RDFHDHGHOD5HJLyQGHO%tR%tR&KLOHFINOT9/ET AL.
*D\DQD%RW
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Chile.
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actualmente casi completamente reemplazado por
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subfamilia Pooideae comprende la mayor parte
HVSHFLHV VHJXLGD SRU 3DQLFRLGHDH
HVSHFLHV&KORULGRLGHDHHVSHFLHV
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Trin. var. KLUWLÀRUXP +DFN /RXLV0DULH
SUHVHQWHV WDPELpQ HQ $UJHQWLQD 3DURGL Helictotrichon bulbosum sólo habita en Chile y
$UJHQWLQD3DURGL\HVHO~QLFRUHSUHVHQWDQWH
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Introducidas
Total
Subfamilia
1ž
1ž
1ž
1ž
Aristidoideae
Arundinoideae
1
1
Bambusoideae
5
Chloridoideae
5
11
Danthonioideae
11
(KUKDUWRLGHDH
1
1
Panicoideae
15
Pooideae
58
171
Total
*D\DQD%RW
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Introducidas
Total
15
51,VOD0RFKD
7
51fXEOH
515DOFR
1
613HQtQVXODGH+XDOSpQ
8
58
61/REHUtDGH&RETXHFXUD
8
'HODVHVSHFLHVLQWURGXFLGDVCynosurus cristatus
/\Corynephorus divaricatus3RXUU%UHLVWUKDQ
sido colectadas hasta ahora sólo en la región del
%tR%tR Corynephorus divaricatusLQWURGXFLGDHQ
&KLOHKDVLGRVHxDODGDVyORSDUDHVWD5HJLyQGHO
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VX LQWURGXFFLyQ HQ 3RU RWUD SDUWH HVSHFLHV
FX\D SUHVHQFLD QR KDEtD VLGR GRFXPHQWDGD HQ
HVWD UHJLyQ GHO SDtV VRQ Bromus madritensis, B.
tectorumFestuca acanthophylla, F. monticola, F.
purpurascensF. tunicata y Rostraria cristata.
ENDEMISMOS
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5HJLyQGHO%tR%tRVHGLVWULEX\HQHQVXEIDPLOLDV
%DPEXVRLGHDH'DQWKRQLRLGHDH\3RRLGHDH'HODV
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WUHVHVSHFLHVHQGpPLFDVGH&KLOHHQOD5HJLyQGHO
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aureofulva y Rytidosperma quirihuense. De la
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GH&KLOH7DEOD,,,(OJpQHURFRQPD\RUQ~PHUR
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N. duriuscula N. gibba N. juncea 3KLO N.
macrathera N. parodii y 1 S¿VWHUL 6yOR FXDGUtFXODVSUHVHQWDURQHVSHFLHVHQGpPLFDV)LJ
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arvensis Anthoxanthum altissimum Bromidium
trisetoidesChusquea quilaDanthonia araucana
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looseriana Gymnachne koelerioides Melica
commersonii M. violacea Nassella duriuscula
Piptochaetium angolenseP. depresum y Polypogon
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distribuidas son Chusquea quila y Melica violacea
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regional son Agrostis arvensis TXH VH FRQRFH
solamente por el material original colectado por
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amethystina y Polypogon linearis son anuales.
DISTRIBUCIÓN ALTITUDINAL
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disminuye consistentemente con la altitud. Tanto
las especies nativas como las introducidas muestran
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chonoticaPoa holciformisP. stenanthaTrisetum
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VREUHORVP
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1 Agrostis arvensis
Agrostis umbellata
Anthoxanthum altissimum
Bromidium trisetoides
5 Chusquea cumingii
Chusquea quila
7 Danthonia araucana
8 Danthonia chilensis var. aureofulva
Deschampsia looseriana
Festuca tunicata
11 Gymnachne koelerioides
Melica commersonii
Melica violacea
Nassella duriuscula
15 Nassella gibba
Nassella juncea
57
17 Nassella macrathera
18 Nassella parodii
1DVVHOODS¿VWHUL
Phalaris amethystina
Piptochaetium angolense
Piptochaetium depressum
Poa cumingii
3RDS¿VWHUL
Polypogon linearis
Rytidosperma quirihuense
1
Trisetum nancaguense
151
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